Nick Upton / Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...
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Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...
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Nick Upton / Bloody-nosed beetles (Timarcha tenebricosa) mating on a grass stem in a chalk grassland meadow, Wiltshire, UK, April.
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Guy Edwardes / 2020VISION / Dawn mist and rolling chalk downland viewed from Wilmington Hill, Wilmington, South Downs National Park, East Sussex, England, UK, July 2011
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Guy Edwardes / 2020VISION / Rolling chalk downland and arable farmland viewed from Wilmington Hill, Wilmington, South Downs National Park, East Sussex, England, UK, July.
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Guy Edwardes / 2020VISION / Wilmington Long Man on Wilmington Hill, a chalk figure cut into the hillside in the 16th century, South Downs National Park, East Sussex, England, UK, July 2011
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Guy Edwardes / 2020VISION / Six-spot burnet moth (Zygaena filipendulae) resting on Red Clover (Trifolium pratense) Kingcombe Meadows DWT reserve, Dorset, UK, May
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Guy Edwardes / 2020VISION / Grass covered anthills in ancient chalk grassland at dawn, Parsonage Down NNR, Wiltshire, UK, June 2011
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Nick Upton / Cherhill white horse, first cut into chalk downland in 1780, Wiltshire, UK, spring 2009
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Nick Upton / Hoary plantain (Plantago media) flowering in a chalk grassland meadow, Wiltshire, UK. June.
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Nick Upton / Hairy violets (Viola hirta) flowering on a chalk grassland slope, Wiltshire, UK. March.
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Nick Upton / Hairy violets (Viola hirta) flowering on a chalk grassland slope, Wiltshire, UK. March.
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Nick Upton / Barren strawberry / Strawberryleaf cinquefoil (Potentilla sterilis) flowering on chalk grassland slope, near Bath, UK. March.
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Nick Upton / Welsh oak longhorn beetle (Pyrrhidium sanguineum) basking on limestone rock in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Male Common fox spider (Alopecosa pulverulenta) crawling over a path in chalk grassland, Marlborough Downs, Wiltshire, UK. May.
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Nick Upton / Dingy skipper (Erynnis tages) butterfly sunning itself in chalk grassland, Morgan's Hill Nature Reserve, Wiltshire, UK. May.
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Nick Upton / Green tiger beetle (Cicindela campestris) hunting on a chalk grassland slope, Wiltshire, UK. March.
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Nick Upton / Grizzled skipper (Pyrgus malvae) butterfly sunning on a Dandelion (Taraxacum officinale) seedhead on chalk grassland, Marlborough Downs, Wiltshire, UK. May.
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Nick Upton / Small yellow underwing (Panemeria tenebrata) moth, nectaring on Germander speedwell (Veronica chamaedrys) flower in chalk grassland, Morgan's Hill, Wiltshire, UK. May.
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Nick Upton / Marbled white butterfly (Melanargia galathea) nectaring on a Small scabious (Scabiosa columbaria) flower on a chalk grassland, Morgan's Hill Reserve, Wiltshire, UK. July.
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Nick Upton / Yellow meadow ant (Lasius flavus) nest mounds on grazed chalk grassland slope, near Swansea, Glamorgan, Wales. July.
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Nick Upton / Mother Shipton moth (Callistege mi) nectaring on Horsehoe vetch (Hippocrepis comosa) flowerhead in a chalk grassland meadow, Wiltshire, UK. June.
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Nick Upton / Rove beetle (Tasgius cf. melanarius) climbing a grass stem in a chalk grassland meadow, Wiltshire, UK. August.
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Nick Upton / Mixed flowers on a chalk grassland down including Round-headed rampion (Phyteuma orbiculare) and Common restharrow (Ononis repens), near Calne, Wiltshire, UK, July.
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Nick Upton / Buff-tailed bumblebee (Bombus terrestris) worker flying to its burrow on chalk grassland hillside with loaded pollen sacs to provision grubs, Marlborough Downs, Wiltshire, UK. May.
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Nick Upton / Solitary mining bee (Andrena sp.) nectaring on Barren strawberry (Potentilla sterilis) flowering on chalk grassland slope, Wiltshire, UK. March.
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars, newly hatched, eating eggshells as others begin spinning a silken larval web over Devil's bit scabious (Succisa pratensis) leaves, their larval food...
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars, newly hatched, spinning a silken larval web over Devil's bit scabious (Succisa pratensis) leaves, their larval food plant, in a chalk grassland mead...
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars, newly hatched, spinning a silken larval web over Devil's bit scabious (Succisa pratensis) leaves, their larval food plant, in a chalk grassland mead...
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars, just hatched, spinning a silken larval web over Devil's bit scabious (Succisa pratensis) leaves, their larval food plant, in a chalk grassland meado...
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars, seven weeks old, feeding on Devil's bit scabious (Succisa pratensis) leaves, their larval food plant, under a silken web they have spun in a chalk g...
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars, three week old, spinning and extending the silken web they have spun over Devil's bit scabious (Succisa pratensis) leaves, their larval food plant,...
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Nick Upton / Seven week old Marsh fritillary (Euphydryas aurinia) caterpillars feeding on Devil's bit scabious (Succisa pratensis) leaves, their larval food plant, under a silken web they have spun in a chalk...
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Nick Upton / Seven week old Marsh fritillary (Euphydryas aurinia) caterpillars feeding on Devil's bit scabious (Succisa pratensis) leaves, their larval food plant, under a silken web they have spun in a chalk...
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Nick Upton / Three week old Marsh fritillary (Euphydryas aurinia) caterpillars spinning and extending the silken web they have spun over Devil's bit scabious (Succisa pratensis) leaves, their larval food plant...
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Nick Upton / Three week old Marsh fritillary (Euphydryas aurinia) caterpillars spinning and extending the silken web they have spun over Devil's bit scabious (Succisa pratensis) leaves, their larval food plant...
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Nick Upton / Marsh fritillary (Euphydryas aurinia) caterpillars hatching and spinning a protective silken larval web on the underside of a Devil's bit scabious (Succisa pratensis) leaf, the larval food plant,...
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Nick Upton / Two-coloured mason bee (Osmia bicolor) turning over a Brown-lipped snail (Cepaeae nemoralis) shell on a chalk grassland slope, Somerset, UK, May. The bee provisions the snail shell with chewed balls o...
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Nick Upton / Two-coloured mason bee (Osmia bicolor) resting at its nest in a Brown-lipped snail (Cepaeae nemoralis) shell before flying away, chalk grassland slope, Somerset, UK, May.
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Nick Upton / Two-coloured mason bee (Osmia bicolor) female flying in to her nest in a Brown-lipped snail (Cepaeae nemoralis) shell to complete sealing up the brood cells inside with chewed up leaves, on a chalk gr...
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Nick Upton / Female Two-coloured mason bee (Osmia bicolor) flying with a long stick to camouflage her nest on a chalk grassland slope, pushing it into position before flying away, Somerset, UK, May.
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Nick Upton / Female Two-coloured mason bee (Osmia bicolor) flying with a long stick to camouflage her nest on a chalk grassland slope, pushing it into position before flying away, Somerset, UK, May.
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Nick Upton / Female Two-coloured mason bee (Osmia bicolor) flying with a long stick to camouflage her nest on a chalk grassland slope, pushing it into position before flying away, Somerset, UK, May.
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Nick Upton / Female Two-coloured mason bee (Osmia bicolor) flying with a long stick to camouflage her nest on a chalk grassland slope, pushing it into position before flying away, Somerset, UK, May.
