Tui De Roy / Epanola saddelback tortoise (Chelonoidis hoodensis), with Galapagos mockingbird (Mimus parvulus) perched on carapace, Espnola Island, Galapagos.

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists, acting as ecosystem engineers by maintaining open meadows and digg...

Tui De Roy / Alcedo giant tortoises (Chelonoidis vandenburghi) group resting in water on moonlit night to keep cool, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) walking, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) pair mating in shallow pool, with two others resting nearby. Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Western Santa Cruz giant tortoises (Chelonoidis porteri) resting in water, South coast (Las Palmitas), Santa Cruz Island, Galapagos

Tui De Roy / Western Santa Cruz giant tortoise (Chelonoidis porteri) feeding on grass, El Chato II, Highlands, Santa Cruz Island, Galapagos

Tui De Roy / Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos

Tui De Roy / Floreana giant tortoise hybrid descendants (Chelonoidis elephantopus) feeding on flower. These are descendants of the extinct Floreana giant tortoise, but are hybridised with other species. Fausto Lle...

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) silhouetted at sunrise, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in pool, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in water at night, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) male mating with female while squabbling with another tortoise. Alcedo Volcano, Isabela Island, Galapagos

Andy Rouse / Alcedo volcano Galapagos giant tortoise (Chelonoidis vandenburghi) adult male on forest track, Urbina Bay, Isabela Island, Galapagos.

Tui De Roy / Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos

Tui De Roy / Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos

Tui De Roy / Hood island giant tortoise (Chelonoidis nigra hoodensis) used for breeding program since 1960s, Tortoise Breeding Centre, Puerto Ayora, Santa Cruz Island, Galapagos Islands, Ecuador, Critically endang...

Tui De Roy / Volcan Alcedo giant tortoises (Chelonoidis nigra vandenburghi) wallowing in rain pools on caldera floor, possibly for thermoregulation or detering parasites, with sulphur fumaroles in background, Alce...

Tui De Roy / Wolf-Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos. Did you know? Galapagos tortoises have a mutualistic relati...

Tui De Roy / Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait, Isabela Island, Galapagos

Pete Oxford / Saddleback form of Galapagos Giant Tortoise (Chelonoidis nigra) walking, Wolf Volcano, Isabela Island, Galapagos

Pete Oxford / Galapagos Giant Tortoise (Geochelone elephantophus / Chelonoidis nigra) head portrait, Charles Darwin Research Station. Puerto Ayora,Santa Cruz Island, Galapagos, endemic

Mark Moffett / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) in Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Pinzon Island Tortoise (Chelonoidis nigra ephippium) pair fighting, Pinzon Island, Galapagos Islands, Ecuador

Tui De Roy / Saddleback Galapagos Tortoise (Chelonoidis hoodensis) in defensive posture, Espanola Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) pair mating, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) on ridge, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) pair in volcanic landscape, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) group in volcanic landscape, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) group wallowing in seasonal pond, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Chatham Island Tortoise (Chelonoidis chathamensis), Galapaguera, San Cristobal Island, Galapagos Islands, Ecuador

Pete Oxford / Indefatigable Island Tortoise (Chelonoidis porteri), Santa Cruz Island, Galapagos Islands, Ecuador

Pete Oxford / Indefatigable Island Tortoise (Chelonoidis porteri) grazing, Santa Cruz Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Ecuador

Pete Oxford / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi), Alcedo Volcano crater floor, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) wallowing in temporary rainy season pool, Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) large male on caldera rim, Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) on caldera rim, Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) juvenile with flowers, Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis vandenburghi) Galapagos Hawk (Buteo galapagoensis) riding on back, Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador