Neil Challis / Giant golden orb weaver spider (Nephila pilipes) focus pull from a small male to much larger female in the background, Western Thailand.
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David Weiller / Aerial tracking shot of locals feeding a juvenile Whale shark (Rhincodon typus) and swimming past a crowd of tourists, Oslob, Philippines, May.
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David Weiller / Aerial tracking shot of locals feeding a juvenile Whale shark (Rhincodon typus) and swimming past a crowd of tourists, Oslob, Philippines, May.
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David Weiller / Aerial tracking shot of coral reef formations in shallow sea, Sipadan, Sabah, Malaysia, November.
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Marie Read / Eastern bluebird (Sialia sialis) male feeding on Winterberries (Ilex verticillata) before flying off, Freeville, New York, USA, January.
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Marie Read / Common loon (Gavia immer) adult swimming with a chick resting on its back, Presque Isle County, Michigan, USA, June.
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Lukas Schaefer / Alpine ibex (Capra ibex) male walking up a mountain in the snow, Gran Paradiso, Italy, December.
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Oscar Dewhurst / European robin (Erithacus rubecula) perched on snow-covered branch, Richmond Park, London, UK. January.
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Jack Dykinga / Yellow-headed blackbird (Xanthocephalus xanthocephalus) flock flying over reed beds, Whitewater Draw Wildlife Preserve, Arizona, USA.
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Robert Thompson / A fallen Sweet chestnut fruit (Castanea sativa) lying on the woodland floor, with a tree trunk covered in Textured lungwort (Lobaria scrobiculata) lichen, UK. October.
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Joel Sartore / Photo Ark / Red-crested turaco (Tauraco erythrolophus) looking around and raising crest, Tracy Aviary. Captive.
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Joel Sartore / Photo Ark / Himalayan monal pheasant (Lophophorus impejanus) male looking around, Pheasant Heaven. Captive.
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Joel Sartore / Photo Ark / Great green macaw (Ara ambiguus) turning its head and moving tongue, Tracy Aviary. Captive.
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Jacob Guy / Scoly coral (Scolymia) feeding under UV light, Corwall, UK. Filmed in studio.
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Jacob Guy / Starry cup coral (Acanthastrea echinata) timelapse of movement under UV light. Filmed in studio.
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Noemie Stroh / Orbicular batfish (Platax orbicularis) schooling into the blue, Shark Reef, Ras Mohamed National park, Red Sea, Egypt, June.
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Noemie Stroh / Yellow scroll coral (Turbinaria reniformis) surrounded by colourful reef fish, Jackson Reef, Red Sea, Egypt, September.
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Matt Roseveare / Parson's chameleon (Calumma parsonii) catching insect with tongue, Eastern rainforests of Madagascar, September.
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Oscar Dewhurst / Mandarin duck (Aix galericulata) drake swimming on pond, backlit by rising sun and bokeh effect, Richmond Park, London, UK. February.
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Bence Mate / Two Great white egret (Ardea alba) in winter, Kiskunsag National Park, Hungary. December.
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Konrad Wothe / Stoat (Mustela erminea) in white winter coat standing upright in snow, Upper Bavaria, Germany, Europe. February.
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Magnus Lundgren / School of Yellow-ribbon sweetlips (Plectorhinchus polytaenia) swimming with the current, Raja Ampat, Indonesia, Pacific Ocean.
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Brian Bevan / Yellowhammer (Emberiza citronella) male singing from bramble then flies away, Bedfordshire, UK, June.
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Bruce Thomson / Crucifix toad / Holy cross frog (Notaden bennetti) sitting in shallow water after heavy summer rain, Westmar, Queensland, Australia.
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Markus Varesvuo / Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October.
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Lewis Jefferies / Close up shot of Pulsating xenid (Heteroxenia fuscescens) feeding , Red Sea, Egypt
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Lewis Jefferies / Common octopus (Octopus vulgaris) changing colour as a threat display, Maldives, Indian Ocean.
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Lewis Jefferies / Powder blue tangs (Acanthurus leucosternon) shoaling, Maldives, Indian Ocean.
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Mateusz Piesiak / Black-headed Gull (Chroicocephalus ridibundus) flock in flight a dawn, Barycz Valley Landscape Park, Poland. Emotion'Ailes Photography Competition 2021 Category Winner - Artistic.
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Mateusz Piesiak / Bramblings (Fringilla montifringilla) feeding on sunflower seeds in a field that could not be cut due to high water levels, Lower Silesia, Poland. Emotion'Ailes Photography Competition 2021, Categ...
Mateusz Piesiak / European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland.
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Guy Edwardes / Ancient Roman bridge and autumn leaves, Allt da Ghob Waterfall, Glen Lyon, Perthshire, Scotland, UK. October.
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Will Burrard-Lucas / Drone shot of Fagradalsfjall Volcano erupting, with lava flow from crater. Iceland, Europe. June 2021.
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David Pattyn / Great white egret (Ardea alba) fishing in shallow pond, with Grey heron (Ardea cinerea) and other Great white egrets in background, Valkenhorst Nature Reserve, The Netherlands, August.
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Jack Dykinga / Ironwood Forest National Monument, Arizona. Sunset light with whispy clouds over Ragged Top Mountain in the Silverbell range.
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Jack Dykinga / Female Burrowing owl (Athene cunicularia) with two chicks in farm fields, near Marana, Arizona, USA.
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Eduardo Blanco / Pale burrower mayfly (Ephoron virgo), swarming in the millions on street at night, appearing like snow. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain, August.
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Eduardo Blanco / Tree silhouetted against volcanic eruption, Cumbre Vieja Volcano, La Palma, Canary Islands, September 2021.
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David Pattyn / Great crested grebe (Podiceps cristatus) with young chicks on the back among Fringed water lilies (Nymphoides peltata) Lake Kerkini, Greece, July.
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Juergen Freund / Bioluminescent fungi (possibly Omphalotus nidiformis / Pleurotus nidiformis) glowing on tree trunk in rainforest at night, Atherton Tablelands, Queensland, Australia. Highly commended in the Plants an...
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Oriol Alamany / Trees silhouetted against erupting Cumbre Vieja volcano at night, La Palma, Canary Islands, September 2021
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Oriol Alamany / Cumbre Vieja volcano erupting at night, La Palma, Canary Islands, September 2021.
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Uri Golman / Close up of a silverback Mountain gorilla (Gorilla beringei beringei) face with eyes closed, and hand / fingers, Virunga National Park, Democratic Republic of Congo.
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Uri Golman / Harp seal (Pagophilus groenlandicus) pup swimming in a hole in the ice, Magdalen Islands, Canada. March.
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Eduardo Blanco / Close up lava from volcanic eruption, Cumbre Vieja Volcano, La Palma, Canary Islands. September 2021
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Eduardo Blanco / Volcanic eruption with silhouette of tree, Cumbre Vieja Volcano, La Palma, Canary Islands. September 2021
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Eduardo Blanco / Volcanic eruption, Cumbre Vieja Volcano, La Palma, Canary Islands. September 2021