Jack Dykinga / Lightning storm with rainbow over Saguaro cactus (Carnegiea gigantea) near Redrock, Arizona State Trust, Sonoran Desert, Arizona. September 2015. Long exposure with lightning trigger.

Klaus Echle / Red Fox (Vulpes vulpes) walking along a fallen trunk, silhouetted in misty forest. Black Forest, Germany, November.

Martin Dohrn / Herd of giraffes (Giraffa camelopardalis) resting at night, Masai Mara, Kenya. Image taken at night using thermal camera technology, with no artificial light.

Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.

Mateusz Piesiak / Black-headed Gull (Chroicocephalus ridibundus) flock in flight a dawn, Barycz Valley Landscape Park, Poland. Emotion'Ailes Photography Competition 2021 Category Winner - Artistic.

Graham Eaton / Barn owl (Tyto alba) flying at night over track between trees, Pwllheli, Wales, UK. April. Runner-up in Nature and Wildlife category of Monivisions Photography Awards 2021.

Juergen Freund / Bioluminescent fungi (possibly Omphalotus nidiformis / Pleurotus nidiformis) glowing on tree trunk in rainforest at night, Atherton Tablelands, Queensland, Australia. Highly commended in the Plants an...

Milan Radisics / Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic volumetric lighting. Vertes Mountains, Hungary.

Guy Edwardes / Woman walking holding umbrella walking through dark woodland near Symondsbury, Bridport, Dorset, England, UK. August 2020

Will Burrard-Lucas / Male melanistic leopard (Panthera pardus) at night, under a starry sky. Laikipia Wilderness Camp, Kenya. Communication Arts Annual 62 Competition 2021 Winner Book Category; Sony World Photographer Awa...

Will Burrard-Lucas / A mating pair of leopards (Panthera pardus) caught in front of a camera trap at night, Laikipia Wilderness Camp, Kenya. Communication Arts Annual 62 Competition 2021 Winner Book Category; Sony World P...

Will Burrard-Lucas / Male melanistic leopard (Panthera pardus) at night, under a starry sky. Laikipia Wilderness Camp, Kenya. Photographed with a camera trap. Nature Photographer of the Year Competition 2021 Winner - Anim...

Guy Edwardes / Fungus (Mycena sp) at night. New Forest National Park, England, UK. October.

Andy Murch / Cape fur seal (Arctocephalus pusillus) playing with Dark shyshark (Haploblepharus pictus) carcass, seal diving in background. False Bay, South Africa. May.

Danny Green / European river otter (Lutra lutra) in mist reflected in water at night. Lincolnshire, UK, September.

Markus Varesvuo / Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February.

Nick Upton / Common bee fly (Bombylius major) nectaring on a Greater periwinkle flower (Vinca major), Wiltshire garden, UK, April.

Shane Gross / Caribbean reef squid (Sepioteuthis sepioidea) portrait at night, Eleuthera, Bahamas.

Shane Gross / Caribbean reef squid (Sepioteuthis sepioidea) portrait reflecting off the surface of the ocean at night, Eleuthera, Bahamas.

Doug Gimesy / Portrait of a Ghost bat (Macroderma gigas) with dark background, captive, Adelaide Zoo, Australia.

Guy Edwardes / Milky Way and Mars over Pondfield Cove, Tyneham, Jurassic Coast World Heritage Site, Dorset, England, UK, August.

Guy Edwardes / Milky Way over Stair Hole, Lulworth, Jurassic Coast World Heritage Site, Dorset, England, UK, August.

Markus Varesvuo / Great grey owl (Strix nebulosa) Kuhmo, Finland, March.

Markus Varesvuo / Great Grey Owl (Strix nebulosa) Kuhmo Finland, March.

Markus Varesvuo / Great Grey Owl (Strix nebulosa) hunting over snow, Kuhmo Finland, March.

Espen Bergersen / Northern fulmar (Fulmarus glacialis) in front of the Seljalandsfoss waterfall, Iceland. May

Andres M. Dominguez / Little owl (Athene noctua) taking shelter from the rain, Arcos de la Frontera, southern Spain. April.

Andres M. Dominguez / Riisitunturi in winter with Aurora Borealis, Kuusamo, Lapland, Finland. February

Alex Mustard / Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies.

Gabriel Rojo / Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina.

Eduardo Blanco / Pale burrower mayfly (Ephoron virgo), swarming in the millions on street at night, appearing like snow. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August 2019. Finalist i...

Eduardo Blanco / European mantis (Mantis religiosa) hunting swarming Pale burrower mayfly (Ephoron virgo), mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August.

Eduardo Blanco / Pale burrower mayfly (Ephoron virgo) swarming around streetlights at night, with gecko (Tarentola mauritanica) inside lamp. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. Aug...

Nigel Cattlin / Immature caterpillar of a codling moth (Cydia pomonell) eating its way through a gallery in a ripening apple fruit, Berkshire, July

Doug Gimesy / Little penguin (Eudyptula minor) on rocks of St Kilda breakwater, Melbourne city lights in background. Victoria, Australia. January 2017.

Doug Gimesy / Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016. COP26 Countdown Photo Competi...

SCOTLAND: The Big Picture / Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK.

Angelo Gandolfi / Forest fire in the Cerrado during dry season. Chapada dos Veadeiros National Park, Goias, Brazil. September 2010.

Paul Hobson / Leaf-nosed bat (Phyllostomidae sp), three nectaring on flower. Costa Rica.

Paul Hobson / Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica.

Paul Hobson / Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October.

Paul Hobson / Badger (Meles meles) crossing road in residential area at night. Sheffield, England, UK. October 2018.

Paul Hobson / Badger (Meles meles) emerging through gateway in wall at night. In urban area, Sheffield, England, UK. October 2018.