Jack Dykinga / Lightning storm with rainbow over Saguaro cactus (Carnegiea gigantea) near Redrock, Arizona State Trust, Sonoran Desert, Arizona. September 2015. Long exposure with lightning trigger.

Klaus Echle / Red Fox (Vulpes vulpes) walking along a fallen trunk, silhouetted in misty forest. Black Forest, Germany, November.

Martin Dohrn / Herd of giraffes (Giraffa camelopardalis) resting at night, Masai Mara, Kenya. Image taken at night using thermal camera technology, with no artificial light.

Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.

Guy Edwardes / Milky Way and Mars over Pondfield Cove, Tyneham, Jurassic Coast World Heritage Site, Dorset, England, UK, August.

Guy Edwardes / Milky Way over Stair Hole, Lulworth, Jurassic Coast World Heritage Site, Dorset, England, UK, August.

Espen Bergersen / Northern fulmar (Fulmarus glacialis) in front of the Seljalandsfoss waterfall, Iceland. May

Andres M. Dominguez / Little owl (Athene noctua) taking shelter from the rain, Arcos de la Frontera, southern Spain. April.

Andres M. Dominguez / Riisitunturi in winter with Aurora Borealis, Kuusamo, Lapland, Finland. February

Alex Mustard / Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies.

Eduardo Blanco / Pale burrower mayfly (Ephoron virgo), swarming in the millions on street at night, appearing like snow. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August 2019. Finalist i...

Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...

Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...

Nigel Cattlin / Immature caterpillar of a codling moth (Cydia pomonell) eating its way through a gallery in a ripening apple fruit, Berkshire, July

Doug Gimesy / Little penguin (Eudyptula minor) on rocks of St Kilda breakwater, Melbourne city lights in background. Victoria, Australia. January 2017.

Doug Gimesy / Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016.

SCOTLAND: The Big Picture / Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK.

Angelo Gandolfi / Forest fire in the Cerrado during dry season. Chapada dos Veadeiros National Park, Goias, Brazil. September 2010.

Paul Hobson / Leaf-nosed bat (Phyllostomidae sp), three nectaring on flower. Costa Rica.

Paul Hobson / Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica.

Paul Hobson / Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October.

Paul Hobson / Badger (Meles meles) crossing road in residential area at night. Sheffield, England, UK. October 2018.

Paul Hobson / Badger (Meles meles) emerging through gateway in wall at night. In urban area, Sheffield, England, UK. October 2018.

Patricio Robles Gil / Kashir grey langur (Semnopithecus ajax) Dachigam National Park, Jammu & Kashmir, India.

Will Burrard-Lucas / Young male melanistic leopard (Panthera pardus), Laikipia Wilderness Camp, Kenya. Photographed with a camera trap.

Will Burrard-Lucas / Young male melanistic leopard (Panthera pardus), Laikipia Wilderness Camp, Kenya. Photographed with a camera trap.

Will Burrard-Lucas / Young male melanistic leopard (Panthera pardus), Laikipia Wilderness Camp, Kenya. Photographed with a camera trap.

Will Burrard-Lucas / Young male melanistic leopard (Panthera pardus), Laikipia Wilderness Camp, Kenya. Photographed with a camera trap.

Will Burrard-Lucas / Young male melanistic leopard (Panthera pardus), Laikipia Wilderness Camp, Kenya. Photographed with a camera trap.

Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.

Sven Zacek / Ural owl (Strix uralensis) perched on Alder (Alnus sp) tree at dusk. Tartumaa, Southern Estonia. April.

Guy Edwardes / Vik i Myrdal church in winter, with full moon, Iceland. January 2015

Guy Edwardes / Northern Lights over Vestrahorn, Hofn, Iceland. September 2017

Guy Edwardes / The Dark Hedges, Stranocum, Co. Antrim, Northern Ireland, September 2010.

Guy Edwardes / Aurora Borealis over Sycamore Gap, Hadrian's Wall, Northumberland, England. February 2014

David Tipling / Barn owl(Tyto alba) flying, North Norfolk, England, UK. february

Ann & Steve Toon / Nile crocodile (Crocodylus niloticus) at night, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Solvin Zankl / Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen and is for mating displays or defenc...

Guy Edwardes / Natural Arch, Siabost, Isle of Lewis, Outer Hebrides, Scotland, UK. March, 2015.

Guy Edwardes / River Etive with isolated cottage beneath Buachaille Etive Mor, Rannoch Moor, Glen Coe, Highlands, Scotland, UK. November, 2013.

Guy Edwardes / Fairy Pools and Black Cuillins, Glen Brittle, Isle of Skye, Inner Hebrides, Scotland, UK. February 2017.

Guy Edwardes / Eilean Glas Lighthouse, Isle of Scalpay, Outer Hebrides, Scotland, UK. March 2015.

Guy Edwardes / Colmer's Hill, Bridport, Dorset, England, UK. December 2016.

Guy Edwardes / Gold Hill in winter, Shaftesbury, Dorset, England, UK. January 2010.

David Allemand / Two Snow geese (Anser caerulescens) during migration, with dark clouds against the sun, Quebec, Canada, October.

Five Films / Timelapse of a lightning storm at night, Murchison Falls National Park, Uganda, April 2014.

Five Films / Timelapse of a lightning storm at night, Murchison Falls National Park, Uganda, April 2014.

Jack Dykinga / Narrows with canyon wall reflections in mud patterned shore, Vermilion Cliffs National Monument, Paria Canyon Wilderness, Arizona, USA.