Nature Production / Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan.
01357049
Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition
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Mark Payne-Gill / Three banded armadillo {Tolypeutes tricinctus} uncurling from defensive ball
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Vishal Jadhav / Exhausted Short-eared owl (Asio flammeus) surrounded by House crow (Corvus splendens) group at dawn, one crow dives down to attack, Maharashtra, India, April.
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Vishal Jadhav / Short-eared owl (Asio flammeus) stood surrounded by a group of House crows (Corvus splendens) trying to find weakness by snapping at its feathers, Maharashtra, India, April.
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Nick Upton / Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK,...
01692243
Matt Roseveare / Caddisfly larva (Trichoptera) building armoured casing / case with pebbles on riverbed, Cornwall, UK, April. Controlled conditions.
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Lewis Jefferies / Common octopus (Octopus vulgaris) changing colour as a threat display, Maldives, Indian Ocean.
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Simon Littlejohn / Common kingfisher (Alcedo atthis) perched outside its nest, opening its wings and beak in an aggressive display to a Common blackbird (Turdus merula) which eventually flies away, River Stour, Dorset,...
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Doug Gimesy / Sub-Antarctic / Brown skua (Stercorarius antarcticus) flying away with egg stolen from a Royal penguin colony (Eudyptes schlegeli), Macquarie Island, Australian Territory. December.
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Staffan Widstrand / Wild Wonders of China / Oriental / Crested honey buzzard (Pernis ptilorhynchus) feeding on honeycomb surrounded by bees, Chayi, Taiwan Controlled conditions
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Staffan Widstrand / Wild Wonders of China / Oriental / Crested honey buzzard (Pernis ptilorhynchus) feeding on honeycomb surrounded by bees, Chayi, Taiwan. Controlled conditions
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Oscar Dewhurst / Western marsh harrier (Circus aeruginosus) being mobbed by black-headed gulls (Chroicocephalus ridibundus) as it takes off with chick. Suffolk, England, UK. June
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Ripan Biswas / Portrait of an ant-mimicking crab spider. (Amyciaea sp) raising its legs to warn the photographer. Buxa tiger reserve, India.
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Stefan Christmann / Emperor penguin (Aptenodytes fosteri) male peering out, looking over the huddle during polar night, Antarctic winter, Atka Bay, Antarctica. July. Bookplate.
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Alex Mustard / Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home, a Haddon's sea anemone (Stichoda...
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Simon Littlejohn / Juvenile Lammergeier vulture (Gypaetus barbatus) in flight, mobbed by a raven, Grand Bargy cirque, Haute Savoie, France, October.
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Simon Littlejohn / Mute swans (Cygnus olor) hissing and lunging at a European otter (Lutra lutra), River Stour, Dorset, England, UK, June.
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Sven Zacek / Ural Owl (strix uralensis) female with talons ready to attack a ringing ornithologist, Tartumaa county, Southern Estonia. May.
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Franco Banfi / Octopus, (Octopus vulgaris), splashing a cloud of ink and swim away with Bearded fireworm, (Hermodice carunculata) who want to eat a small octopus, top of the wall of Bisevo, Vis Island, Croatia, Adri...
01639937
Dave Bevan / Slow motion clip of a Great spotted woodpecker (Dendrocopos major) returning to nest hole, attacked by a Nuthatch (Sitta europaea), Carmarthenshire, Wales, UK, April.
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Tony Heald / African buffalo (Syncerus caffer) herd showing defensive behaviour as an African lioness (Panthera leo) walks by, Mana Pools National Park, Zimbabwe. September....
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Tony Heald / Lion (Panthera leo) moving away from a defensive African buffalo (Syncerus caffer) Mana Pools National Park, Zimbabwe.
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Matthew Maran / Red fox (Vulpes vulpes) cub defending its kill,a large Brown rat (Rattus norvegicus) from another cub, North London, UK. August. Highly commended in the Behaviour : Mammals Category of the Wildlife Ph...
01631790
Morley Read / Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador.
Nigel Cattlin / Large rose sawfly (Arge pagana) larvae feeding on ornamental rose leaves in summer, Berkshire, September
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John Shaw / Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June.
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Espen Bergersen / Aerial shot of a female Humpback whale (Megaptera novaeangliae) and calf surrounded by Killer whales (Orcinus orca), mother hits sideways with her tail fluke to drive Killer whales away, Skjervoy, Tro...
01620835
Konrad Wothe / Larvae of Birch sawfly (Craesus septentrionalis) in alarm position, Upper Bavaria, Germany. October.
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John Chan / Western gulls (Larus occidentalis) attacking an Elegant tern (Thalasseus elegans) with a broken wing, Southern California, USA, April.
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Tony Wu / Sardines (Clupeidae) fleeing from predators in the early morning. They were being herded and chased by Trevallies and Blacktip reef sharks in the water, and picked off by brown boobies and terns from...
01618684
Klein & Hubert / Common hedgehog (Erinaceus europaeus) young orphan, curled in a ball in grass. France.
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Klein & Hubert / Common hedgehog (Erinaceus europaeus) hoglet aged five days, eyes not yet open
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Klein & Hubert / Common hedgehog (Erinaceus europaeus) hoglet age one week, curled into a ball, France.
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Klein & Hubert / Hedgehog (Erinaceus europaeus) female sleeping with her hoglets, age 6 days, France.
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Klein & Hubert / Hedgehog (Erinaceus europaeus) carrying hoglet, age 10 days to new nest. France.
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Klein & Hubert / Hedgehog (Erinaceus europaeus) three hoglets, age three wee weeks, following adult female. France.
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Klein & Hubert / Hedgehog (Erinaceus europaeus) hoglets age six days, France.
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Klein & Hubert / Hedgehog (Erinaceus europaeus) hoglets uncurling from ball, age 10 days, France.
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Georgette Douwma / Boxer Crab (Lybia tessellata) with sea anemones in its claws for defense. Lembeh Strait, North Sulawesi, Indonesia.
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Guy Edwardes / Mozambique spitting cobra (Naja mossambica) spitting venom. Kruger, South Africa. Controlled conditions.
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Robert Valentic / Eastern tiger snake (Notechis scutatus) female, Yarra River corridor in suburban Melbourne, Victoria, Australia, in mid-summer. She is in a defensive pre-strike position with hood expanded, accompanie...
01607921
Konrad Wothe / Slow motion clip of Wood ants (Formica rufa) on anthill, squirting formic acid to repel threat, Bavaria, Germany, April.
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Five Films / Female Grey seal (Halichoerus grypus) defending pup on beach, chasing male away, Horsey, Norfolk, November.
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Lucas Bustamante / Tropical shield mantis (Choeradodis rhombicollis) in defensive position, on flower, Chinambi, Carchi, Ecuador.
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Remi Masson / Group of Black bullhead (Ameiurus melas) shoaling to protect themselves against predators, River Rhone, Rhone-Alpes, France, July.
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Robert Thompson / Sawfly larvae (Nematus pavidus) defensive posture, Peatlands Park, County Armagh, Northern Ireland. September.