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    Owen Newman / Male Red kangaroos fighting {Macropus rufus} Sturt NP, New South Wales Australia
    21100942
    Owen Newman / Male Red kangaroos fighting {Macropus rufus} Sturt NP, New South Wales Australia
    Sylvain Cordier / RF-Gelada baboon (Theropithecus gelada) dominant male running. Debre Libanos, Rift Valley, Ethiopia. 2017. (This image may be licensed either as rights managed or royalty free)
    01643101
    Sylvain Cordier / RF-Gelada baboon (Theropithecus gelada) dominant male running. Debre Libanos, Rift Valley, Ethiopia. 2017. (This image may be licensed either as rights managed or royalty free)
    01625450
    John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...
    Klein & Hubert / Meerkat (Suricata suricatta) alpha female nursing her one month old pups on the den, Kalahari Desert, Botswana.
    01605869
    Klein & Hubert / Meerkat (Suricata suricatta) alpha female nursing her one month old pups on the den, Kalahari Desert, Botswana.
    Carol Walker / Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA.
    01546952
    Carol Walker / Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA.
    Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) courtship dance, Cardiff, UK, March.
    01514654
    Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) courtship dance, Cardiff, UK, March.
    Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
    01514090
    Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
    Cyril Ruoso / Bonobo (Pan paniscus) male and female copulating, Lola ya Bonobo Sanctuary, Democratic Republic of Congo
    01508672
    Cyril Ruoso / Bonobo (Pan paniscus) male and female copulating, Lola ya Bonobo Sanctuary, Democratic Republic of Congo
    Ole Jorgen Liodden / Waved albatross (Phoebastria irrorata) pair in courtship display, Espanola, Galapagos Islands. Critically endangered species.
    01507396
    Ole Jorgen Liodden / Waved albatross (Phoebastria irrorata) pair in courtship display, Espanola, Galapagos Islands. Critically endangered species.
    Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cubs with mother 'T19 Krishna' in background, Ranthambhore National Park, India.
    01501088
    Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cubs with mother 'T19 Krishna' in background, Ranthambhore National Park, India.
    Markus Varesvuo / Black grouse (Lyrurus tetrix) mating in snow, Utajarvi, Finland, May.
    01498210
    Markus Varesvuo / Black grouse (Lyrurus tetrix) mating in snow, Utajarvi, Finland, May.
    David Pattyn / Great crested grebe (Podiceps cristatus) pair performing their 'weed dance' in which they take a vertical position in front of each other and offer plant material to each other. The Netherlands. March...
    01492184
    David Pattyn / Great crested grebe (Podiceps cristatus) pair performing their 'weed dance' in which they take a vertical position in front of each other and offer plant material to each other. The Netherlands. March...
    Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.
    01428042
    Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.
    Denis-Huot,Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.
    01427795
    Denis-Huot,Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.
    01423099
    Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.
    Anup Shah / Topi (Damaliscus lunatus jimela) two males sparring, with another approaching, Masai Mara National Reserve, Kenya. March
    01412867
    Anup Shah / Topi (Damaliscus lunatus jimela) two males sparring, with another approaching, Masai Mara National Reserve, Kenya. March
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while his juvenile son 'Tembo' aged 4 years waits his turn, Bai Ho...
    01406120
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while his juvenile son 'Tembo' aged 4 years waits his turn, Bai Ho...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while female 'Malui' and their juvenile son 'Tembo' aged 4 years s...
    01406112
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while female 'Malui' and their juvenile son 'Tembo' aged 4 years s...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while female 'Malui' and their juvenile son 'Tembo' aged 4 years s...
    01406110
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while female 'Malui' and their juvenile son 'Tembo' aged 4 years s...
    David Pattyn / Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
    01405454
    David Pattyn / Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
    David Pattyn / Red necked grebe (Podiceps grisegena) approaching a competitor in an attempt to drive him away during the courting season, Danube area, Bulgaria, April
    01405413
    David Pattyn / Red necked grebe (Podiceps grisegena) approaching a competitor in an attempt to drive him away during the courting season, Danube area, Bulgaria, April
    Anup Shah / African elephants (Loxodonta africana) sparring, Masai Mara National Reserve, Kenya, August
    01402482
    Anup Shah / African elephants (Loxodonta africana) sparring, Masai Mara National Reserve, Kenya, August
    Charlie Summers / Plains Zebra (Equus quagga) in a dominance fight with dust flying, Lake Magadi, Serengeti, Tanzania.
    01399906
    Charlie Summers / Plains Zebra (Equus quagga) in a dominance fight with dust flying, Lake Magadi, Serengeti, Tanzania.
    Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
    01396877
    Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
    Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
    01396874
    Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
    Carol Walker / Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010
    01393067
    Carol Walker / Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010
    George Sanker / Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
    01388526
    George Sanker / Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
    Rolf Nussbaumer / Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...
    01388033
    Rolf Nussbaumer / Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...
    Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October. Did you know? Richmond Park deer herds are descended from historic medieval royal hunting stock.
    01378127
    Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October. Did you know? Richmond Park deer herds are descended from historic medieval royal hunting stock.
    Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October
    01378126
    Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October
    Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, with blocks of flats in the background, Richmond Park, London, England, UK, October
    01378125
    Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, with blocks of flats in the background, Richmond Park, London, England, UK, October
    Marie Read / Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
    01377664
    Marie Read / Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
    Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on and a Herring Gull (Larus argentus) flies past. Scandinavia, April.
    01377568
    Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on and a Herring Gull (Larus argentus) flies past. Scandinavia, April.
    Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on. Scandinavia, April. 5th, 2010 Euronatur Wildlife photographer competition.
    01377567
    Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on. Scandinavia, April. 5th, 2010 Euronatur Wildlife photographer competition.
    Gerrit Vyn / Dall sheep (Ovis dalli) rams head butting, one behaviour by which males establish dominance during the Autumn mating season. Denali National Park, Alaska, USA
    01377428
    Gerrit Vyn / Dall sheep (Ovis dalli) rams head butting, one behaviour by which males establish dominance during the Autumn mating season. Denali National Park, Alaska, USA
    Andy Rouse / Two Common Zebra (Equus quagga) fighting. Masai Mara, Kenya.
    01374825
    Andy Rouse / Two Common Zebra (Equus quagga) fighting. Masai Mara, Kenya.
    Terry Whittaker / 2020VISION / Red deer (Cervus elaphus) stag chasing another during a fight, rutting season, Bushy Park, London, UK, October
    01371861
    Terry Whittaker / 2020VISION / Red deer (Cervus elaphus) stag chasing another during a fight, rutting season, Bushy Park, London, UK, October
    Terry Whittaker / 2020VISION / Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October
    01371859
    Terry Whittaker / 2020VISION / Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October
    Terry Whittaker / 2020VISION / Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October (This image may be licensed either as rights managed or royalty free.)
    01371857
    Terry Whittaker / 2020VISION / Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October (This image may be licensed either as rights managed or royalty free.)
    Terry Whittaker / 2020VISION / Two Fallow deer (Dama dama) bucks during fight, rutting season, Richmond Park, London, UK, October (This image may be licensed either as rights managed or royalty free.)
    01371835
    Terry Whittaker / 2020VISION / Two Fallow deer (Dama dama) bucks during fight, rutting season, Richmond Park, London, UK, October (This image may be licensed either as rights managed or royalty free.)
    Markus Varesvuo / Horned / Slavonian grebe (Podiceps auritus) two individuals fighting in water, Finland May
    01368856
    Markus Varesvuo / Horned / Slavonian grebe (Podiceps auritus) two individuals fighting in water, Finland May
    Terry Whittaker / 2020VISION / Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
    01359732
    Terry Whittaker / 2020VISION / Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
    Terry Whittaker / 2020VISION / Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
    01359731
    Terry Whittaker / 2020VISION / Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
    Terry Whittaker / 2020VISION / Konik horse (Equus caballus) herd grazing with two stallions interacting, Wicken Fen, Cambridgeshire, UK, June 2011
    01359729
    Terry Whittaker / 2020VISION / Konik horse (Equus caballus) herd grazing with two stallions interacting, Wicken Fen, Cambridgeshire, UK, June 2011
    Radomir Jakubowski / Two Muskox (Ovibos moschatus) in landscape, Dovrefjell national park, Norway, September
    01357948
    Radomir Jakubowski / Two Muskox (Ovibos moschatus) in landscape, Dovrefjell national park, Norway, September
    Anup Shah / Adolescent male Lions (Panthera leo) play fighting, Masai Mara National Reserve, Kenya, March
    01356398
    Anup Shah / Adolescent male Lions (Panthera leo) play fighting, Masai Mara National Reserve, Kenya, March
    Dave Watts / Eastern grey kangaroo (Macropus giganteus) two males boxing, Australian Capital Territory, Australia, December
    01354800
    Dave Watts / Eastern grey kangaroo (Macropus giganteus) two males boxing, Australian Capital Territory, Australia, December
    Andrew Parkinson / Whooper swan (Cygnus cygnus) adult aggressively chasing another, UK visitor between November and March. Lancashire, UK, October.
    01354353
    Andrew Parkinson / Whooper swan (Cygnus cygnus) adult aggressively chasing another, UK visitor between November and March. Lancashire, UK, October.
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