Owen Newman / Male Red kangaroos fighting {Macropus rufus} Sturt NP, New South Wales Australia
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Laurie Hedges / Bison (Bison bison) bull kicking up dust, urinating and then rolling on the ground for scent, Montana, USA, August.
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Sylvain Cordier / RF-Gelada baboon (Theropithecus gelada) dominant male running. Debre Libanos, Rift Valley, Ethiopia. 2017. (This image may be licensed either as rights managed or royalty free)
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Sylvain Cordier / Gelada baboon (Theropithecus gelada) female carrying baby on back, dominant male in background. Debre Libanos, Rift Valley, Ethiopia.
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Alex Mustard / Western clown anemonefish (Amphiprion ocellaris) poses with a large, dominant female behind. The clownfish in the foreground is probably a juvenile. Bitung, North Sulawesi, Indonesia. Lembeh Strait, M...
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)
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John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) dominant male (T29) cooling off in a waterhole Kanha National Park, Central India. Camera trap image.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) dominant male (T29) cooling off in a waterhole Kanha National Park, Central India. Camera trap image.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) dominant male (T29) and resident female (T27) cooling off in a watering hole Kanha National Park, Central India. Camera trap image.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) dominant male (T29) and resident female (T27) cooling off in a waterhole Kanha National Park, Central India. Camera trap image.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) female (T27) acting cautiously as dominant male (T29) approaches watering hole Kanha National Park, Central India. Camera trap image.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) dominant male (T29) walking on mud dam wall. Kanha National Park, Central India. Camera trap image.
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Yashpal Rathore / Bengal tiger (Panthera tigris tigris) dominant male approaching 'Beja pani' waterhole, Kanha National Park, Central India. Camera trap image.
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Christophe Courteau / Mountain gorilla (Gorilla beringei beringei), dominant silverback
in forest clearing. Volcanoes National Park, Rwanda.
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Klein & Hubert / Meerkat (Suricata suricatta) alpha female nursing her one month old pups on the den, Kalahari Desert, Botswana.
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Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) courtship dance, Cardiff, UK, March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Fred Olivier / Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia.
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Cyril Ruoso / Bonobo (Pan paniscus) male and female copulating, Lola ya Bonobo Sanctuary, Democratic Republic of Congo
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Ole Jorgen Liodden / Waved albatross (Phoebastria irrorata) pair in courtship display, Espanola, Galapagos Islands. Critically endangered species.
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Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cubs with mother 'T19 Krishna' in background, Ranthambhore National Park, India.
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Markus Varesvuo / Black grouse (Lyrurus tetrix) mating in snow, Utajarvi, Finland, May.
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David Pattyn / Great crested grebe (Podiceps cristatus) pair performing their 'weed dance' in which they take a vertical position in front of each other and offer plant material to each other. The Netherlands. March...
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Jabruson Motion / Small group of African forest elephants (Loxodonta africana cuclotis) running into forest clearing, trumpeting to dominant male feeding and cooling down in a mineral pool, Dzanga Bai, Dzanga-Ndoki Nat...
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Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.
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Denis-Huot,Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.
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Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.
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Anup Shah / Topi (Damaliscus lunatus jimela) two males sparring, with another approaching, Masai Mara National Reserve, Kenya. March
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Fiona Rogers / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African R...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years peering through vegetation, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged about 32 years scratching his head, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Rep...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) rear view of dominant male Silverback 'Makumba' aged approx 32 years standing on a forest track, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while his juvenile son 'Tembo' aged 4 years waits his turn, Bai Ho...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years standing portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. D...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while female 'Malui' and their juvenile son 'Tembo' aged 4 years s...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting in a forest clearing, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...
01406110
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while female 'Malui' and their juvenile son 'Tembo' aged 4 years s...
01406109
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while his juvenile son 'Tembo' aged 4 years waits his turn, Bai Ho...
01406108
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
01406107
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
01406100
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
01406099
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
01406098
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting in a forest clearing, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years walking through Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dece...
01406085
Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years walking through the forest, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river while sub-adult female 'Mosoko' aged 8 years crosses bi-pedally, Bai Hokou, Dza...
