Lewis Jefferies / Motion timelpase of river and sun setting through trees at Kennall Vale nature reserve, Ponsanooth, Cornwall, UK, January.
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Richard Hopkins / Slow motion shot of Short-eared owl (Asio flameus) taking off from the ground and flying out of frame, Gloucestershire, UK, March.
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Neil Aldridge / Timelapse of the sun rising over the Okavango Delta in northern Botswana.
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John Chan / American avocets (Recurvirostra americana) foraging in a tidal basin, Bolsa Chica Ecological Reserve, Southern California, USA, October.
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John Chan / American white pelicans (Pelecanus erythrorhynchos) foraging in tidal basin at sunset, Bolsa Chica Ecological Reserve, Southern California, USA, September.
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Matthew Maran / Red fox cubs (Vulpes vulpes) play fighting in an allotment, London, England, UK, June.
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Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July
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Ismaele Tortella / Four Eurasian elk (Alces alces) walking through vegetation , Norway, August
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Ismaele Tortella / Aerial tracking shot over Tromso in the winter, northern Norway, March 2019.
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Fred Olivier / Aerial shot of Castleton Tower at sunset, Castle Valley, Moab, Utah, USA, 2018.
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Fred Olivier / Alkefjellet cliffs at dusk, with Brunnich's guillemot (Uria lomvia) breeding colony, Alkefjellet, Svalbard, Norway, 2016.
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Morley Read / Reventador Volcano erupting at dawn, Ecuador, 2017. The mountain has been in a constant state of eruption since 2002.
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Morley Read / Aerial shot ascending over rainforest canopy at dawn, with the Rio Napo in background, a tributary of the Amazon, Napo Province, Ecuador, 2017.
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Milan Radisics / Aerial shot of Gunnuhver steam geyser, Gunnhuver Hot Springs, Reykjanes Peninsula, Iceland, August 2018.
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Milan Radisics / Aerial shot of rootless cones or pseudocraters, formed by steam explosions as flowing hot lava crosses over a wet surface, South Iceland, August 2018.
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John Chan / Great horned owl (Bubo virginianus) chicks stretching and flapping their wings, Southern California, USA, May.
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John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019
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Espen Bergersen / Aerial shot of a Killer whale (Orcinus orca) spyhopping several times close to land, with people watching and taking pictures, Sommaroy, Troms. Norway, January.
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Espen Bergersen / Aerial shot of a Humpback whale (Megaptera novaeangliae) diving and showing tail fluke, Troms, Norway, January.
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Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) spyhopping, with a snowy landscape in the background, Skjervoy, Troms, Norway, December.
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Espen Bergersen / Aerial shot of a small pod of Killer whales (Orcinus orca) at surface, with snowy landscape in the background, Skjervoy, Troms, Norway, December.
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John Waters / Large murmuration of Common starlings (Sturnus vulgaris) swirling over reedbed roost at sunset, Ham Wall RSPB Reserve, Somerset, England, UK, January.
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John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.
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John Waters / Large flock of Common starlings (Sturnus vulgaris) swirling over reedbed roost at dusk, Ham Wall RSPB Reserve, Somerset, England, UK, November.
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Nick Upton / Two Red fox (Vulpes vulpes) cubs playing near the entrance to their den at dusk, Somerset, England, UK, May.
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Nick Upton / Male Eurasian beaver (Castor fiber) grazing at dusk, River Otter, Devon, England, UK, June.
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Juergen Freund / Timelapse from day to night looking over Tsing Yi container port, with cranes filling container ships, Hong Kong, China, 2017.
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Juergen Freund / Timelapse of Fireflies (Lampyridae) flying around a tree, Bicol, Philippines.
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Juergen Freund / Timelapse of Fireflies (Lampyridae) flying around a tree, Bicol, Philippines.
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Juergen Freund / Timelapse from day to night of bioluminescent fungi (Omphalotus nidiformis / Pleurotus nidiformis) on a tree trunk, Atherton Tablelands, Queensland, Australia.
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Juergen Freund / Timelapse from day to night looking over Tsing Yi container port, with cranes filling container ships, Hong Kong, China, 2017.
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Juergen Freund / Little red flying foxes (Pteropus scapulatus) flying from roosting tree at dusk, Atherton Tablelands, Queensland, Australia.
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Juergen Freund / Australian freshwater crocodiles (Crocodylus johnsoni) catching a Little red flying fox (Pteropus scapulatus) flying to drink from a watering hole, Kimberley, Western Australia.
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Juergen Freund / Australian freshwater crocodiles (Crocodylus johnsoni) trying to catch Little red flying foxes (Pteropus scapulatus) flying to drink from a watering hole, Kimberley, Western Australia
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Juergen Freund / Large group of Little red flying foxes (Pteropus scapulatus) flying at dusk, Atherton Tablelands, Queensland, Australia.
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Juergen Freund / Large group of Little red flying foxes (Pteropus scapulatus) flying at dusk, Atherton Tablelands, Queensland, Australia.
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Gavin Hellier / Aerial shot tracking along chalk cliffs at sunset, near Old Harry Rocks, Dorset, England, UK, May 2017. Hellier
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Gavin Hellier / Timelapse of clouds moving over Bamburgh Castle at sunset, Northumberland, England, UK, October 2017. Hellier
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Gavin Hellier / Timelapse of the view from the summit of Pao de Acucar or Sugarloaf Mountain at sunset, Rio de Janeiro, Brazil, South America - 4K time lapseSeptember 2016. Hellier
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Gavin Hellier / Iguacu Falls, Foz do Igua?u, Iguacu (Iguazu) National Park, Brazil, South AmericaSeptember 2016. Hellier
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Gavin Hellier / Timelapse of beachgoers on Ipanema Beach at sunset, with Dois Irmaos mountain in the background, Rio de Janeiro, Brazil, September 2016. Hellier
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Gavin Hellier / Timelapse from day to night looking over Botafogo bay, with Pao de Acucar or Sugarloaf mountain in the background, Rio de Janeiro, Brazil, South America - 4K time lapseSeptember 2016. Hellier
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Gavin Hellier / Timelapse from day to night looking over the Gaudi House Museum, Barcelona, Catalonia, Spain, May 2016. Hellier
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Gavin Hellier / Timelapse looking out of the front of a train on the elevated Dubai Metro System, Dubai, United Arab EmiratesJanuary 2017. Hellier
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Gavin Hellier / Timelapse from day to night looking towards the Peter and Paul Fortress, St, Petersburg, Russia, May 2016. Hellier
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Gavin Hellier / Timelapse from day to night of people visiting the illuminated ice sculptures at the Harbin Ice and Snow Festival, Heilongjiang Province, China, February 2015. Hellier
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John Waters / Wide-angle shot of wetland landscape at sunset, with Lapwings (Vanellus vanellus) flying to roost, Ham Wall RSPB Reserve, Somerset Levels, England, UK, February.
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Five Films / Large flock of Common starlings (Sturnus vulgaris) flying at dusk, Somerset Levels, England, UK, November.