Jan Wegener / Common Nighthawk (Chordeiles minor) male in display flight, Winnie, Texas
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Marie Read / Ruby-throated hummingbird (Archilochus colubris) male flies in to feed on nectar from Eastern red columbine flower (Aquilegia canadensis), Freeville, New York, USA, June.
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John Waters / Golden plover (Pluvialis apricaria) flock taking off and flying over saltmarsh, Steart Marshes WWT Reserve, Somerset, UK, November.
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Jack Dykinga / Yellow-headed blackbird (Xanthocephalus xanthocephalus) flock flying over reed beds, Whitewater Draw Wildlife Preserve, Arizona, USA.
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Vishal Jadhav / Red-breasted flycatcher (Ficedula parva) male hunting flying insects from a rock, Maharashtra, India, February.
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Nick Upton / Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK,...
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Vishal Jadhav / Asian green bee-eater (Merops orientalis) hunting insects from perch in the grass, Maharashtra, India, July.
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Oliver Wright / Marbled white butterfly (Melanargia galathea) sequence showing landing and taking off from a Greater knapweed (Centaurea scabiosa) flower. Yorkshire, UK. Digital composite.
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Oliver Wright / A female kingfisher (Alcedo atthis) fishing through an ice hole in winter, emerging with prey in beak. Leeds, Yorkshire, UK. January.
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Oliver Wright / A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January.
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Mateusz Piesiak / Black-headed Gull (Chroicocephalus ridibundus) flock in flight a dawn, Barycz Valley Landscape Park, Poland. Emotion'Ailes Photography Competition 2021 Category Winner - Artistic.
Graham Eaton / Barn owl (Tyto alba) flying at night over track between trees, Pwllheli, Wales, UK. April. Runner-up in Nature and Wildlife category of Monivisions Photography Awards 2021.
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Doug Gimesy / Greater crested tern (Thalasseus bergii) flies over breaking waves on a stormy day, with a rainbow in the background, Apollo Bay, Victoria, Australia.
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Simon Littlejohn / Common kingfisher (Alcedo atthis) perched outside its nest, opening its wings and beak in an aggressive display to a Common blackbird (Turdus merula) which eventually flies away, River Stour, Dorset,...
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Marie Read / Front view of an adult Black skimmer (Rynchops niger) in flight carrying a fish, Long Island, New York, USA. August.
Awarded a Judges Choice in the Birds category in the 2020 North American Nature As...
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Kim Taylor / Male Cockchafer beetle (Melolontha melolontha) unfolding its wings before taking off to fly away, Surrey, UK, May.
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Richard Hopkins / Slow motion shot of Short-eared owl (Asio flameus) taking off from the ground and flying out of frame, Gloucestershire, UK, March.
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Richard Hopkins / Slow motion shot of Short-eared owl (Asio flameus) flying over grassland searching for prey, before it dives into the grass and then flies away, Gloucestershire, UK, March.
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Oscar Dewhurst / Shelduck (Tadorna tadorna) landing on water. Gloucestershire, England, UK. March
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Neil Aldridge / Slow motion shot of Osprey (Pandion haliaetus) diving for and flying off with fish, Rothiemurchus, Cairngorms National Park, Scotland, UK, July.
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John Cancalosi / Gila Woodpecker (Melanerpes uropygialis) bringing food to nest in Saguaro cactus (Carnegiea gigantea) before leaving, Sonoran Desert, Arizona, USA.
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Doug Gimesy / Sub-Antarctic / Brown skua (Stercorarius antarcticus) flying away with egg stolen from a Royal penguin colony (Eudyptes schlegeli), Macquarie Island, Australian Territory. December.
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Jack Dykinga / Sandhill cranes (Antigone candensis), in flight against the setting full moon at dawn. Whitwater Draw, Arizona Game and Fish Wildlife area, Arizona, USA.
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John Chan / Calliope hummingbird (Stellula calliope) chasing insects before returning to its perch, Southern California, USA, April.
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John Chan / Monarch butterfly (Danaus plexippus) taking flight from a Pride of madeira (Echium candicans) after contact with a Honey bee (Apis mellifera), Southern California, USA, April.
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John Chan / Elegant terns (Thalasseus elegans) gathering at mussel bed during breeding season, one hovering tern has a mussel clamped to its foot, a common mishap caused by stepping on an open mussel, Bolsa Chica...
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Neil Aldridge / Slow motion shot of two Grey long eared bat (Plecotus austriacus) flying, leaving roost in barn to go hunting, Devon, UK. The Grey long eared bat is one of the UK's rarest mammals.
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Neil Aldridge / Slow motion shot of Soprano pipistrelle bat (Pipistrellus pygmaeus) crawling out of roost in barn roof and flying away, Devon, UK.
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Kim Taylor / Barn swallow (Hirundo rustica), flying over a cattle trough, descending vertically to come in low over the water and drink. Slow motion. The rapid passing of the swallow causes Water boatmen / Backswi...
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Kim Taylor / Female Red mason bee (Osmia bicornis) returning to her nest hole in a dead holly trunk, April.
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Tony Wu / Japanese dwarf flying squirrel (Pteromys volans orii) gliding through snowy forest, Hokkaido, Japan.
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Doug Gimesy / White-capped mollymawk / albatross (Thalassarche cauta) in flight, Foveaux Strait, Stewart Island, New Zealand. Winner of Animal Portrait category in the Australian Geographic Nature Photographer of...
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Brian Bevan / Male Kingfisher (Alcedo atthis) landing on branch after capturing fish then uses perch to stun prey before flying away, Bedfordshire, UK, July.
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Brian Bevan / Wood pigeon (Columba palumbus) resting with chick in nest before catching a fly and eating it, Bedfordshire, UK, August.
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Markus Varesvuo / Arctic tern (Sterna paradisaea) in flight over water, multiple reflections in sea. Iceland. June.
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Stephen Dalton / Indian leaf butterfly (Kallima paralekta) in flight.
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Stephen Dalton / Barn owl (Tyto alba) in flight approaching bell-tower, carrying rodent prey to young in nest. Sussex, UK.
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Oscar Dewhurst / Canada geese (Branta canadensis) in flight low over lake at sunrise in mist. London, UK. November
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Stephen Dalton / Orange tip butterfly (Anthocharis cardamines) male in flight amongst Lady's smock or Cuckooflower (Cardamine pratensis),England.
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Stephen Dalton / Black witch moth (Ascalapha odorata) in flight, Venezuela. This large moth has a 7 inch wingspan. Wingscales being shed during flight.
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Sven Zacek / Brimstone butterfly(Gonepteryx rhamni) male and female during mating flight, early spring. Tartumaa county, Southern Estonia. May.
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Ernie Janes / Small white butterfly (Pieris rapae) and Small tortoiseshell butterfly (Aglais urticae) in flight, Norfolk, England, UK, June.
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David Tipling / Red grouse (Lagopus lagopus) male in flight over Heather. Grinton Moor, above Swaledale, Yorkshire Dales National Park, England, UK. July.
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David Tipling / Red grouse (Lagopus lagopus) male in flight. Grinton Moor, above Swaledale, Yorkshire Dales National Park, England, UK. July.
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David Tipling / Flock of Knot (Calidris canutus) chased by a Peregrine as they arrive to roost at high tide, Cormorants in foreground, Snettisham RSPB Reserve, The Wash, Norfolk, September.
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Andres M. Dominguez / Crested tit (Lophophanes cristatus) about to land on Elmleaf blackberry (Rubus ulmifolius) plant, Sierra de Grazalema Natural Park, Southern Spain, January.
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Jeff Foott / California condor (Gymnogyps californianus) landing, forest in background. Bird with tags on wings. Near San Pedro Martir National Park, Northern Baja California, Mexico. 2017.