Graham Eaton / Barn owl (Tyto alba) flying at night over track between trees, Pwllheli, Wales, UK. April. Runner-up in Nature and Wildlife category of Monivisions Photography Awards 2021.

Alex Mustard / Oceanic whitetip shark (Carcharhinus longimanus) is framed by the surface. Elphinstone reef, Marsa Alam, Egypt. Red Sea.

Neil Aldridge / Steenbok (Raphicerus campestris) female standing framed by a fallen tree, Mapungubwe National Park, Limpopo Province, South Africa.

Yashpal Rathore / Demoiselle crane (Anthropoides virgo) flying silhouetted and framed by Jarokha (carved windows) of a Haveli or townhouse. Khichan, Western Rajasthan, India. December.

Lucas Bustamante / Anthony's poison-frog (Epipedobates anthonyi) seen through hole in leaf. Buenaventura Reserve, El Oro, Ecuador.

Pete Oxford / San Lucas marsupial frog (Gastrotheca pseustes) captive, occurs in Ecuadorian Andes.

Erlend Haarberg / Frost covered European beech tree (Fagus sylvatica) in a snow covered field framed by frosty branches, Calonda, Harghita County, Transylvania, Romania, January.

Cyril Ruoso / Atlantic puffin (Larus tridactyla) seen from burrow, Iceland, July.

Ole Jorgen Liodden / Boy on skis going up slope, framed by snow laden tree, Norway, February 2014.

Ole Jorgen Liodden / Ivory gull (Pagophila eburnea) behind Polar bear (Ursus maritimus) framed by its legs, Svalbard, Norway, July.

Jack Dykinga / Frost damaged Saguaro cactus (Carnegiea gigantea) with sunset light shining through limbs. South Maricopa Mountains Wilderness, Arizona, USA, March 2015.

Jack Dykinga / Saguaro cactus (Carnegiea gigantea) with twisted arms at sunset, with Ajo Mountains in the background, Arizona, USA.

Tim Laman / Rainforest interior views, looking up.

Alex Mustard / Trees and sky photographed at dawn through the surface of Three Sisters Spring, Crystal River, Florida, United States of America.

Ben Hall / Northern gannets (Morus / Sula bassanus) portrait of individual with a courting pair in foreground, UK, Bass Rock, Scotland, Portrait category winner in the British Wildlife Photography Awards BWPA co...

David Fleetham / Galapagos Sea Lion (Zalophus californianus wollebacki) framed by a school of Black Striped Salema (Xenocys jessiae). Santa Cruz Island, Galapagos Islands, Equador.

Alex Mustard / White's / Golden Seahorse (Hippocampus whitei) framed by marine plants. Chowder Bay, Sydney Harbour, New South Wales, Australia, February.

Edwin Giesbers / Proboscis Monkey (Nasalis larvatus) female sitting in tree, partially framed by soft focus leaves in foreground, Bako National Park, Sarawak, Borneo, Malaysia

Andrew Parkinson / Ptarmigan (Lagopus mutus) in white winter plumage framed by a walker passing by during a blizzard. Cairngorm Mountains, Scotland, UK. February (non-ex)

David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.

David Fleetham / Schooling Raccoon butterflyfish (Chaetodon lunula) framed in a lava formation off the island of Lanai, Hawaii.

Wild Wonders of Europe / Möllers / Fallow deer (Dama dama) buck in wood, framed by two trees, Klampenborg Dyrehaven, Denmark, October 2008

Anup Shah / African lion cubs (Panthera leo) framed by body of lioness, Serengeti NP, Tanzania

Suzi Eszterhas / 8 week Cheetah {Acinonyx jubatus} cubs framed by mother's legs, Masai Mara, Kenya.

Suzi Eszterhas / 8-week Cheetah {Acinonyx jubatus} cubs framed by mother's legs, Masai Mara, Kenya.

Andrew Parkinson / Springbok {Antidorcas marsupialis} framed by Elephant at waterhole. Etosha NP, Namibia

SCOTLAND: The Big Picture / A hillwalker framed by a 'fog bow', on snow-covered Slioch mountain summit, Torridon, Highlands, Scotland, UK. November, 2019.

Colin Varndell / Adult house mice (Mus musculus) posing in mossy square hole. Dorset, UK. March. Captive.

Ross Hoddinott / Banded demoiselle (Calopteryx splendens) male silhouetted on a fern, Lower Tamar Lakes, Cornwall, UK. July.

Ole Jorgen Liodden / Split level view of female mallard (Anas platyrhynchos) resting on the edge of a semi-frozen lake in spring, Norway.

John Waters / Red Campion (Silene dioica) flowers framed by out of focus flowers to create soft foreground, Mendips, UK, July.

Jack Dykinga / Giant sequoia (Sequoiadendron giganteum) tree trunks with White firs (Abies concolor) in background, framed by blooming Western dogwood (Cornus nuttallii), Sequoia National Park, California, USA.

Maxime Aliaga / Water channel with mangroves either side, Komodo National Park, Indonesia.

Ross Hoddinott / RF - Red deer (Cervus elaphus) stag standing amongst Scot's pine (Pinus sylvestris) tree trunks in forest. Cairngorms National Park, Scotland, UK. January. (This image may be licensed either as ri...

Enrique Lopez-Tapia / Sydney Opera House and skyline, viewed at dusk under Sydney Harbour Bridge. New South Wales, Australia. 2008.

Enrique Lopez-Tapia / Granitic boulders at Karlu Karlu / Devils Marbles Conservation Reserve, framed by Eucalyptus tree trunks. Northern Territory, Australia. 2008.

Fabian Muhlberger / Wild Wonders of China / Taiwan yellow tit ( Machlolophus holsti ) framed by leaves. Highlands of Taiwan. Endemic.

Fabian Muhlberger / Wild Wonders of China / Taiwan yellow tit ( Machlolophus holsti ) framed by leaves, Taiwan. Endemic.

Fabian Muhlberger / Wild Wonders of China / Taiwan yellow tit ( Machlolophus holsti ) framed by leaves. Highlands of Taiwan, Taiwan. Endemic.

Fabian Muhlberger / Wild Wonders of China / Taiwan barbet ( Psilopogon nuchalis ) , framed by foliage, in a park in Taipei, Taiwan. Endemic.

Bruno D'Amicis / Cultivated fields and herder with zebu cattle, viewed from church forest. Church forests remain largely intact within a degraded landscape as they are considered sacred. Bahir Dar, Ethiopia. 2018.

Ashley Cooper / Stalactites on ceiling of Milodon Cave, view out of cave to trees and mountains. Milodon Cave Natural Monument, Magallanes, Chile. January 2020.

Edwin Giesbers / Barn owl (Tyto alba) sleeping, portrait. Captive.

Edwin Giesbers / Barn owl (Tyto alba) portrait. Captive.

Juergen Freund / Lava tubes, view out of cave towards forest. Undara Volcanic National Park, Undara is an Aboriginal word translating to 'long way'. Queensland, Australia. 2017.

Jack Dykinga / Chain cholla (Cylindropuntia fulgida) in Sonoran Desert in evening light, Saguaro (Carnegiea gigantea) and Ragged Top Mountain in background. Ironwood National Monument, Arizona, USA. March 2020.

Espen Bergersen / People framed in crystal clear ice on diamond beach, Iceland. Commended in the Nordic Nature Photo Contest 2019.

Philippe Clement / Tre Cime di Lavaredo / Drei Zinnen, three mountain peaks viewed from WW1 cave shelter. Sexten Dolomites seen from WW1 cave shelter, South Tyrol, Italy, October 2019