Sergey Gorshkov / Minden / Northern Goshawk (Accipiter gentilis) feeding on Black Grouse (Tetrao tetrix) female in winter, Finland

Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Utajarvi, Finland, April.

David Tipling / Red grouse (Lagopus lagopus) male in flight over Heather. Grinton Moor, above Swaledale, Yorkshire Dales National Park, England, UK. July.

David Tipling / Red grouse (Lagopus lagopus) male in flight. Grinton Moor, above Swaledale, Yorkshire Dales National Park, England, UK. July.

Markus Varesvuo / Black Grouse (Lyrurus tetrix) male displaying with bokeh effect in the foreground, Liminka, Finland, March

Markus Varesvuo / Black Grouse (Lyrurus tetrix) male displaying with bokeh effect in the foreground, Liminka, Finland, March

Jack Dykinga / Yellow-headed blackbird (Xanthocephalus xanthocephalus) flock murmuration as they take off from the marsh at dawn, Whitewater Draw, Arizona State Game and Fish Reserve, USA. January.

Sven Zacek / Black Grouse (Tetrao tetrix) two males facing off on a lekking site, Varumaa county, Southern Estonia. April.

Andy Sands / Woodcock (Scolopax rusticola) female walking along ditch at edge of road, Upper Teesdale, Co Durham, England, UK, May

Erlend Haarberg / Rock ptarmigan (Lagopus muta) in flight, Tysfjorden, Nordland, Norway, March.

Markus Varesvuo / Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March.

SCOTLAND: The Big Picture / RF - Black Grouse (Tetrao tetrix) male peforming flutter jump display on lek, Cairngorms National Park, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)

SCOTLAND: The Big Picture / RF - Black Grouse (Tetrao tetrix), two males fighting on lek , Scotland, UK.April (This image may be licensed either as rights managed or royalty free.)

Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA

Sven Zacek / Ural owl (Strix uralensis) female on the nest with nestling hidden under wing. Tartumaa, Southern Estonia. April.

Ann & Steve Toon / Spotted eagle owl (Bubo africanus) subadult at nigh by water, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Andy Sands / Grey partridge (Perdix perdix) three birds on snow covered mound, Bedfordshire, England, UK, December

Loic Poidevin / Red-legged partridge (Alectoris rufa) with chicks, Vendee, France, June

Ernie Janes / Pheasant (Phasianus colchicus) male flying to roost at sunset with trees silhouetted in background England, UK. December.

Staffan Widstrand / Wild Wonders of China / Golden pheasant (Chrysolophus pictus) male jumping, Yangxian Biosphere Reserve, Shaanxi, China

Guy Edwardes / Saddle-billed stork (Ephippiorhynchus senegalensis), Moremi Game Reserve, Botswana.

Sergey Gorshkov / Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April

Konrad Wothe / Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June.

Staffan Widstrand / Wild Wonders of China / Golden pheasant (Chrysolophus pictus) male displaying in grass in Yangxian Nature Reserve, Shaanxi, China, September.

Staffan Widstrand / Wild Wonders of China / Red jungle fowl (Gallus gallus) Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April.

Ann & Steve Toon / Pied crows (Corvus albus) perched on horns of antelope skull. Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / African fish eagle (Haliaeetus vocifer) flying, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Egyptian goose (Alopochen aegyptiaca) with goslings, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Espen Bergersen / Ptarmigans (Lagopus mutus) in white winter plumage, camouflaged against mountain scree with snow, Spitsbergen, Svalbard, Norway, April

Espen Bergersen / Ptarmigans (Lagopus mutus) in white winter plumage, camouflaged against mountain scree with snow, Spitsbergen, Svalbard, Norway, April. Highly commended in the Animals in their Environment Category of...

Neil Aldridge / Lesser masked weaver (Ploceus intermedius) male hanging upside down from the entrance, Northern Tuli Game Reserve, Botswana.

Wim van den Heever / Egytian Goose (Alopochen aegyptiaca) rear view bathing, sings spread, Zimanga Private Game Reserves, South Africa

Lou Coetzer / Pintailed whydah (Vidua macroura) male displaying to female, Masai Mara, Kenya.

David Pattyn / Martial eagle (Polemaetus bellicosus) perched on a branch. Masai Mara National Reserve, Kenya, Africa, August.

Markus Varesvuo / Female Capercaillie (Tetrao urogallus) with newly hatched chick on nest, Kuhmo, Finland, June.

Markus Varesvuo / Female Capercaillie (Tetrao urogallus) ready to mate, Vaala, Finland, May.

Markus Varesvuo / Willow grouse / Ptarmigan (Lagopus lagopus) pecking twig, Utsjoki, Finland, April.

Markus Varesvuo / Female Black grouse (Tetrao / Lyrurus tetrix) perched in tree covered in snow, Inari Kiilopaa, Finland, January.

Markus Varesvuo / Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.

Markus Varesvuo / Female Black grouse (Tetrao / Lyrurus tetrix) perched in Willow feeding, Utajarvi, Finland, May.

Markus Varesvuo / Four Grey partridges (Perdix perdix) on snow, Kauhajoki, Finland, January.

Markus Varesvuo / Willow grouse / Ptarmigan (Lagopus lagopus) fluffed up perched in snow, Inari, Finland, February.

Markus Varesvuo / Male Capercaillie (Tetrao urogallus) flying, Jalasjarvi, Finland, April.