Winfried Wisniewski / African Lion (Panthera leo) female nuzzling a male, Masai Mara, Kenya

Carol Walker / Andalusian horsea, two stallions coming together in arena one dappled grey and one pure white. Spain.

David Pattyn / Great crested grebe (Podiceps cristatus) adults courting and greeting each other to confirm their bond. Even with the young chicks already out they often show courtship behavior to each other, Valkenh...

Klein & Hubert / Grey wolves (Canis lupus) pair and juvenile greeting each other, USA. Controlled conditions

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults with chicks aged six to eight weeks. Atka Bay, Antarctica. September.

George Sanker / Mallard (Anas platyrhynchos) pair greeting on snow-covered pond. Acadia National Park, Maine, USA.

George Sanker / Mallard (Anas platyrhynchos)pair greeting on snow-covered pond. Acadia National Park, Maine, USA.

Edwin Giesbers / RF - European ground squirrels / Sousliks (Spermophilus citellus)greeting, Gerasdorf, Austria. April. (This image may be licensed either as rights managed or royalty free.)

Mark Taylor / Yellow Labrador Retriever puppy, 8 weeks old, sitting with raised paw

Ben Hall / Greenfinch (Carduelis chloris) pair, one perched on branch and one hovering in snowfall, Derbyshire, February.

Bence Mate / Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January. Winner of the Portfolio category of the Terre Sauvage Na...

Pal Hermansen / European moose (Alces alces) bulls near houses, Nordland, Norway. July.

Will Burrard-Lucas / Ethiopian wolves (Canis simensis) individual from the BBC pack greeting each other after returning from hunting, Ethiopia.

Tui De Roy / Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Ernie Janes,Ernie Janes / Newly hatched ducklings with eggs.

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) two individuals greeting, Tangkoko National Park, Sulawesi, Indonesia.

Kristel Richard / Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June

Carol Walker / Two white horses of the Camargue, interacting, greeting head to head in marshes, Camargue, Southern France

Andy Rouse / African Lion (Panthera leo) male lion greeting female and cubs, Notch from Disney African Cats film, Masai Mara GR, Kenya, February

Markus Varesvuo / Common Crossbill (Loxia curvirostra) male and female greeting, Kuusamo, Finland, February.

Staffan Widstrand / Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January

Ann & Steve Toon / Two Cape Ground squirrels (Xerus inauris) greeting, Kgalagadi Transfrontier Park, South Africa Non-ex.

Wild Wonders of Europe / Popp-Ha / Matterhorn (4,478m) with reflection in Lake Riffel at sunrise, Switzerland, September 2008. WWE BOOK. WWE OUTDOOR EXHIBITION. NOT TO BE USED FOR GREETING CARDS OR CALENDARS TILL 2013 PRESS IMAGE.

Anup Shah / Spotted hyena pups greeting an adult (Crocuta crocuta). Masai Mara National Reserve, Kenya. December

Wild Wonders of Europe / Hamblin / Konik horse and young foal interacting, Oostvaardersplassen, Netherland, June 2009

Wild Wonders of Europe / Unterthiner / Two European bison (Bison bonasus) fighting, Bialowieza NP, Poland, February 2009. WWE BOOK. WWE OUTDOOR EXHIBITION. NOT TO BE USED FOR GREETING CARDS OR CALENDARS PRESS IMAGE. Wild Wonders kids book.

Andrew Parkinson / Two Kittiwakes (Rissa tridactyla) greeting one another after one returns to the nest site, Farne Islands, UK

Fiona Rogers / Ring-tailed Lemur (Lemur catta) male greeting infant. Berenty Private Reserve, Madagascar. Oct 2008.

Niall Benvie / Moorish gecko {Tarentola mauritanica} juvenile, Spain WWE BOOK. WWE OUTDOOR EXHIBITION. NOT TO BE USED FOR GREETING CARDS OR CALENDARS Wild Wonders kids book.

Niall Benvie / European scorpion {Buthus occitanus} Spain WWE BOOK. WWE OUTDOOR EXHIBITION. NOT TO BE USED FOR GREETING CARDS OR CALENDARS

Eric Baccega / Two semi wild Przewalski horses interacting, greeting (Equus ferus przewalskii), Parc du Villaret, Causse Mejean, Lozere, France

Carol Walker / Woman face to face with Bay Oldenburg filly (Equus caballus) Fort Collins, Colorado. Model released.

Anup Shah / Simien jackal / Ethiopian wolf {Canis simensis} mother greeting cubs at den, Bale Mountains, Bale NP, Ethiopia

Anup Shah / Simien jackals / Ethiopian wolf {Canis simensis} greeting behaviour showing body language, Bale Mountains, Bale NP, Ethiopia

Lynn M Stone / Arctic foxes greeting {Vulpes lagopus} Captive.

Anup Shah / Common zebras greeting {Equus quagga} Masai Mara GR, Kenya East Africa

Richard Du Toit / Spotted hyaenas greeting at dusk {Crocuta crocuta} Okavango delta, Botswana

Anup Shah / Subadult Lions affectionate greeting {Panthera leo} Masai Mara NR Kenya, East Africa

Anup Shah / Khur / Indian wild ass greeting {Equus hemionus khur} Rann of Kutch, Gujarat, India

Anup Shah / Khur / Indian wild ass greeting {Equus hemionus khur} Rann of Kutch, Gujarat, India

Tony Heald / Black backed jackal greeting pup {Canis mesomelas} Etosha NP, Namibia

Anup Shah / Common zebras greeting {Equus quagga} Masai Mara, Kenya East Africa

Anup Shah / Spotted hyaena {Crocuta crocuta} greeting ritual, Masai Mara, Kenya East-Africa