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      • hanifaru
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    Doug Perrine / Researcher Guy Stevens photographs belly spot identification pattern of Reef manta ray (Manta alfredi formerly Manta birostris) feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, Oc...
    01349192
    Doug Perrine / Researcher Guy Stevens photographs belly spot identification pattern of Reef manta ray (Manta alfredi formerly Manta birostris) feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, Oc...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) barrel roll filter feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, October
    01349177
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) barrel roll filter feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain-feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, October
    01349175
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain-feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) vortex / cyclone feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    01349164
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) vortex / cyclone feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Ind...
    01349163
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Ind...
    Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315951
    Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in very shallow water, viewed from the surface. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315950
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in very shallow water, viewed from the surface. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Manta ray (Manta birostris) feeding on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315942
    Alex Mustard / Manta ray (Manta birostris) feeding on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Tui De Roy / Manta Ray (Manta birostris) filter feeding, Hanifaru, Baa Atoll, Maldives
    00598989
    Tui De Roy / Manta Ray (Manta birostris) filter feeding, Hanifaru, Baa Atoll, Maldives
    Tui De Roy / Manta Ray (Manta birostris) filter feeding, Hanifaru, Baa Atoll, Maldives
    00598988
    Tui De Roy / Manta Ray (Manta birostris) filter feeding, Hanifaru, Baa Atoll, Maldives
    Alex Mustard / Manta ray (Mobula alfredi) with mouth open, feeding on plankton near the surface, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean.
    01700581
    Alex Mustard / Manta ray (Mobula alfredi) with mouth open, feeding on plankton near the surface, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean.
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton Hanifaru Bay, Raa Atoll, Maldives
    01642594
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton Hanifaru Bay, Raa Atoll, Maldives
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    01642593
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    01642592
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    01642591
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    01642590
    Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
    Tui De Roy / Reef manta ray (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
    01642589
    Tui De Roy / Reef manta ray (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
    Tui De Roy / Reef manta rays (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
    01642588
    Tui De Roy / Reef manta rays (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) filter feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    01349194
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) filter feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    01349190
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) feeding on plankton at surface, Kihaadufaru Reef, with Dhonfanu Island in background, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, Octob...
    01349188
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) feeding on plankton at surface, Kihaadufaru Reef, with Dhonfanu Island in background, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, Octob...
    Doug Perrine / close up of gills of Reef manta ray (Manta alfredi formerly Manta birostris) feeding, with open gill slits showing gill plates through which plankton is filtered (and which also draw oxygen from the w...
    01349187
    Doug Perrine / close up of gills of Reef manta ray (Manta alfredi formerly Manta birostris) feeding, with open gill slits showing gill plates through which plankton is filtered (and which also draw oxygen from the w...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) with shark bite out of pectoral fin, among group feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, October
    01349186
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) with shark bite out of pectoral fin, among group feeding on plankton, Hanifaru Bay, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / RF- Reef manta ray (Manta birostris / alfredi) feeding on plankton, with unique pattern of underside spots, allowing individual identification, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indi...
    01349185
    Doug Perrine / RF- Reef manta ray (Manta birostris / alfredi) feeding on plankton, with unique pattern of underside spots, allowing individual identification, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indi...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) with cephalic fins rolled up, being cleaned by Moon wrasses (Thalassoma lunare) and Blue-streak cleaner wrasses (Labroides dimidiatus) Hanifaru...
    01349184
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) with cephalic fins rolled up, being cleaned by Moon wrasses (Thalassoma lunare) and Blue-streak cleaner wrasses (Labroides dimidiatus) Hanifaru...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) being cleaned by Moon wrasses (Thalassoma lunare) and Blue-streak cleaner wrasses (Labroides dimidiatus) at mouth of Hanifaru Bay, Hanifaru Lago...
    01349183
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) being cleaned by Moon wrasses (Thalassoma lunare) and Blue-streak cleaner wrasses (Labroides dimidiatus) at mouth of Hanifaru Bay, Hanifaru Lago...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) swimming over cleaning station on patch reef with Yellow / Golden sweepers (Parapriacanthus ransonetti) and other schooling fish, Hanifaru Bay e...
    01349182
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) swimming over cleaning station on patch reef with Yellow / Golden sweepers (Parapriacanthus ransonetti) and other schooling fish, Hanifaru Bay e...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) being cleaned by Moon wrasses (Thalassoma lunare) and Blue-streak cleaner wrasses (Labroides dimidiatus) Hanifaru Bay entrance, Hanifaru Lagoon,...
    01349181
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) being cleaned by Moon wrasses (Thalassoma lunare) and Blue-streak cleaner wrasses (Labroides dimidiatus) Hanifaru Bay entrance, Hanifaru Lagoon,...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) barrel roll filter feeding on plankton, accompanied by juvenile Golden pilot jacks (Gnathanodon speciosus), Hanifaru Bay, Baa Atoll, Maldives, I...
    01349176
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) barrel roll filter feeding on plankton, accompanied by juvenile Golden pilot jacks (Gnathanodon speciosus), Hanifaru Bay, Baa Atoll, Maldives, I...
    Doug Perrine / Diver photographing Reef manta rays (Manta alfredi formerly Manta birostris) at reef cleaning station with cleaner wrasse at mouth of Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean,...
    01349174
    Doug Perrine / Diver photographing Reef manta rays (Manta alfredi formerly Manta birostris) at reef cleaning station with cleaner wrasse at mouth of Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean,...
    Doug Perrine / Reef manta rays(Manta alfredi formerly Manta birostris) filter feeding on plankton, with tourist snorkeling and diving down to touch a ray, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian O...
    01349173
    Doug Perrine / Reef manta rays(Manta alfredi formerly Manta birostris) filter feeding on plankton, with tourist snorkeling and diving down to touch a ray, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian O...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, with tourist snorkeling and diving down to touch a ray, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian...
    01349172
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, with tourist snorkeling and diving down to touch a ray, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, with tourist snorkeling in background and trying to touch a ray, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives...
    01349171
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, with tourist snorkeling in background and trying to touch a ray, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives...
    Doug Perrine / (Manta alfredi formerly Manta birostris) filter feeding on plankton, one with a lot of white dorsal pigmentation, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    01349170
    Doug Perrine / (Manta alfredi formerly Manta birostris) filter feeding on plankton, one with a lot of white dorsal pigmentation, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    01349169
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, with snorkelers and scuba divers in background, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, O...
    01349168
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton, with snorkelers and scuba divers in background, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, O...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among school of Fusilier fish (Caesionidae), Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, Octob...
    01349167
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among school of Fusilier fish (Caesionidae), Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, Octob...
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indi...
    01349166
    Doug Perrine / Reef manta ray (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indi...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Ind...
    01349165
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Ind...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain filter feeding on plankton, swimming over school of Fusilier fish (Caesionidae) Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, India...
    01349162
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain filter feeding on plankton, swimming over school of Fusilier fish (Caesionidae) Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, India...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain filter feeding on plankton among Silverside fish in feeding aggregation, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean...
    01349161
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain filter feeding on plankton among Silverside fish in feeding aggregation, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean...
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain-feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October, Digitally enhanced image.
    01349160
    Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) chain-feeding on plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean, October, Digitally enhanced image.
    Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315953
    Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315952
    Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in very shallow water, viewed from the surface. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315949
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in very shallow water, viewed from the surface. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in very shallow water. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315948
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in very shallow water. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in a very shallow water. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315947
    Alex Mustard / Large Manta ray (Manta birostris) feeding on plankton in a very shallow water. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    Alex Mustard / Manta ray (Manta birostris) in shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
    01315944
    Alex Mustard / Manta ray (Manta birostris) in shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.
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