John Waters / Golden plover (Pluvialis apricaria) flock taking off and flying over saltmarsh, Steart Marshes WWT Reserve, Somerset, UK, November.
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Doug Perrine / Schooling female Scalloped hammerhead shark (Sphyrna lewini), Big Island, Hawaii, Pacific Ocean.
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Oscar Dewhurst / Red knot (Calidris canutus) flock at high tide roost. Snettisham, Norfolk, UK. October.
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David Pattyn / Common starling (Sturnus vulgaris) murmuration in between high voltage powerlines and electricity pylons, The Netherlands, Europe. February.
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David Pattyn / Common starling (Sturnus vulgaris) flock gathering in their thousands at their roosting place, De Houtwiel Nature Reserve, The Netherlands, Europe. February.
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David Pattyn / Common starling (Sturnus vulgaris) murmuration in between high voltage powerlines and electricity pylons, The Netherlands, Europe. February.
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Eduardo Blanco / Pale burrower mayfly (Ephoron virgo), swarming in the millions on street at night, appearing like snow. Mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain, August.
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Ben Cranke / King penguins (Aptenodytes patagonicus) congregate on the beach, seeking a safe point to go into the sea, with male Southern elephant seals (Mirounga leonina) blocking their path. St Andrew's Bay,...
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Doug Gimesy / Sub-Antarctic / Brown skua (Stercorarius antarcticus) flying away with egg stolen from a Royal penguin colony (Eudyptes schlegeli), Macquarie Island, Australian Territory. December.
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Nick Upton / Common carder bumblebee (Bombus pascuorum) nectaring on a Dandelion (Taraxacum officinale) flowerhead among many Common daisies (Bellis perennis) on a lawn left unmown to allow wild flowers to bloom t...
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John Chan / Elegant terns (Thalasseus elegans) gathering at mussel bed during breeding season, one hovering tern has a mussel clamped to its foot, a common mishap caused by stepping on an open mussel, Bolsa Chica...
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Theo Bosboom / Long line of walking people with headlights, heading back at night from visiting Fagradalsfjall volcano on Iceland, photographed around half an hour after sunset. The eruption draws large numbers of v...
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Theo Bosboom / Group of tourists silhouetted against volcanic eruption, gathered at one of the viewing points close to the craters of the Fagradalsfjall volcano on Iceland. The best viewing points differ from day to...
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Theo Bosboom / Long line of people, mainly local, walking with headlights, heading back from a visit to the Fagradalsfjall volcano, Iceland. Long exposure showing direction of movement. 2 April 2021
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Claudio Contreras / Caribbean flamingos (Phoenicopterus ruber) adults and chicks in breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
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Pal Hermansen / Flock of domestic geese moving together in a field, Vestfold, Norway.
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Staffan Widstrand / Wild Wonders of China / Oriental / Crested honey buzzard (Pernis ptilorhynchus) feeding on honeycomb surrounded by bees, Chayi, Taiwan Controlled conditions
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Staffan Widstrand / Wild Wonders of China / Oriental / Crested honey buzzard (Pernis ptilorhynchus) feeding on honeycomb surrounded by bees, Chayi, Taiwan. Controlled conditions
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David Pattyn / Common starling (Sturnus vulgaris) murmuration, flock gathering above trees before landing at winter roost. The Netherlands. February.
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David Pattyn / Common starling (Sturnus vulgaris) murmuration, flock pursued by Peregrine falcon (Falco peregrinus) before landing at winter roost. The Netherlands. February 2020.
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David Pattyn / Common starling (Sturnus vulgaris) murmuration, flocks gathering above trees before landing at winter roost. The Netherlands. February 2020.
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David Pattyn / Common starling (Sturnus vulgaris) murmuration, flock gathering in butterfly shape before landing at winter roost. The Netherlands. February.
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David Pattyn / Common starling (Sturnus vulgaris) murmuration, flock gathering above trees before landing at winter roost. The Netherlands. February.
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Lucas Bustamante / Monarch butterflies (Danaus plexippus) flying during a warm morning. When the overwintering period draws to an end large numbers become active. Michoacan, Mexico, February.
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Philippe Clement / Wood anemones (Anemone nemorosa) in flower in forest, dolly shot, Belgium, April.
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Carol Walker / Mustang herd standing in snow. Black Hills Wild Horse Sanctuary, South Dakota, USA. January 2017.
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Simon Littlejohn / Flock of Griffon vultures (Gyps fulvus) feeding, Andalusia, Spain, May.
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Brandon Cole / Golden jellyfish (Mastigias papua),Jellyfish Lake, Palau.
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Brandon Cole / Golden jellyfish (Mastigias papua), Jellyfish Lake, Palau.
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Brandon Cole / Golden jellyfish (Mastigias papua), Jellyfish Lake, Palau.
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Brandon Cole / Diver with Golden jellyfish (Mastigias papua), Jellyfish Lake, Palau. Model released.
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Mark Carwardine / Aerial shot zooming in on a superpod of Short-beaked common dolphins (Delphinus delphis), Sea of Cortez, Baja California, Mexico.
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Bence Mate / Dying Common frogs (Rana temporaria) with their legs removed for food, left to die in their breeding pool surrounded by frogspawn. Covasna, Romania. Winner of Nature category in World Press Photo awar...
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Erlend Haarberg / Reindeer herders on snowmobiles, moving a large flock of semi-domesticated Reindeer (Rangifer tarandus), to the reindeer calving areas in the Jotunheimen National Park, Norway. April.
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Orsolya Haarberg / Reindeer herders moving a large herd of semi-domesticated Reindeer (Rangifer tarandus) with the help of snowmobiles, to the reindeer calving areas in the Jotunheimen National Park, Norway. April.
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Orsolya Haarberg / Reindeer herders moving a large flock of semi-domesticated Reindeer (Rangifer tarandus), with the help of snowmobiles, to the reindeer calving areas in the Jotunheimen National Park, Norway. Here, the...
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David Hall / Split level of Sockeye salmon (Oncorhynchus nerka) during migration; Adams River, British Columbia, Canada. October.
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Danny Green / Starling murmuration, hundreds of thousands of starlings (Sturnus vulgaris) gather in the skies near their night time roost. Gretna Green, Scotland, January. Black and White Category Winner, Wildlife...
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John Cancalosi / Juvenile Timber rattlesnakes (Crotalus horridus) massed together at nest site, Pennsylvania, USA.
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John Cancalosi / Timber rattlesnakes (Crotalus horridus) at nest site with young, Pennsylvania, USA.
Doug Gimesy / During an extreme heat-stress event at Melbourne's Yarra Bend Grey-headed Flying-fox (Pteropus poliocephalus) colony, where temperatures exceeded 43 degreesC, in a desperate search for somewhere coole...
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Ismaele Tortella / Eurasian otter (Lutra lutra) hunting in a shoal of Atlantic herring (Clupea harengus), northern Norway, March
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Ismaele Tortella / Northern gannet (Morus bassanus) colony, chick climbing back into nest, with the midnight sun, Norway, July
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Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.
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Fred Olivier / Clark's anemonefish (Amphiprion clarkii) and Three spot damselfish (Dascyllus trimaculatus) swimming around anemone (Stichodactyla), Nusatupe Lagoon, Solomon Islands.
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Fred Olivier / Tracking shot of a Brunnich's guillemot (Uria lomvia) breeding colony, nesting on cliff ledges, Alkefjellet, Svalbard, Norway.
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Fred Olivier / Brunnich's guillemot (Uria lomvia) nesting on cliff ledges, Alkefjellet, Svalbard, Norway.