Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.

Mark MacEwen / King penguin (Aptenodytes patagonicus) group walking past Southern elephant seal (Mirounga leonina) colony, mountains in background. St Andrews Bay, South Georgia. October 2017.

Nick Garbutt / RF - Ring-tailed lemurs (Lemur catta) back lit and sun bathing at first light. Berenty Private Reserve, southern Madagascar. (This image may be licensed either as rights managed or royalty free.)

Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December

Alex Hyde / Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November.

Daniel Heuclin / Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018.

Edwin Giesbers / RF - Banded darter (Sympetrum pedemontanum) group of five resting, Netherlands, September. (This image may be licensed either as rights managed or royalty free.)

Yashpal Rathore / Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December.

Tony Wu / Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March.

Doug Perrine / False killer whale (Pseudorca crassidens) pod, off the Kona Coast of Hawaii Island, Hawaiian Islands, USA. January.

Bence Mate / Great egret (Ardea alba) group of five at water's edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February.

Bence Mate / Group of five Brown-hooded parrots (Pyrilia haematotis) in rain, Costa Rica.

Wim van den Heever / Blue wildebeest (Connochaetes taurinus) herd silhouetted against the rising sun with clouds in the background. Greater Kruger National Park, South Africa

Wim van den Heever / Plains zebra (Equus quagga) group of four including foal at sunset, Savuti Marsh, Botswana.

Jeff Vanuga / White horses of the Camargue galloping through marshes in the Camargue, France. April.

Jordi Chias / Bryde's whale (Balaenoptera edeni) and Common dolphins (Delphinus delphis) chasing small fish offshore. Santa Maria Island, Azores, Atlantic Ocean. Third Place in the Underwater Category of the Por El...

Jabruson / Mozambican children with captive Yellow Baboon youngster (Papio cynocephalus) young animal caught during troop crop raiding. Pemba to Montepuez highway, north-eastern Mozambique, November 2011. Specia...

Pal Hermansen / Inuit sled dogs resting, near Ittoqortoormiit, Scoresbysund, Greenland. Second Prize in the Nature category of the Spider Black and White Photography Competition 2015.

Staffan Widstrand / Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa. Specially commended in the Animal Portraits category of...

Klein & Hubert / Meerkat (Suricata surricatta) group of five warming up in the sunshine, Kalahari desert, South Africa.

Alex Mustard / Group of predators hunting together, in an example of 'Nuclear foraging'. The Moray eel (Gymnothorax javanicus) flushes out prey, and is followed by a Napolean wrasse (Cheilinus undulatus) and a Bluef...

Bence Mate / Great egrets (Ardea alba) and Grey heron (Ardea cinerea) squabbling over fish, with some in flight. Blurred motion photograph, Lake Csaj, Pusztaszer, Hungary. February.

Pascal Pittorino / Corsican pig (Sus scrofa domestica) sow, suckling the piglets, Parc Naturel Regional de Corse / Natural Regional Park of Corsica, Corsica Island, France. September.

Cyril Ruoso / Ringed-tailed lemur (Lemur catta) foraging in sisal field looking for flowers during a poor year without much fruit. Berenty reserve, Madagascar, September.

Cyril Ruoso / Keeper feeding Common woolly monkeys (Lagothrix lagotricha) in Ikamaperou Sanctuary, Amazon, Peru. October 2006.

Cyril Ruoso / Vicuna (Vicugna vicugna) herd in Altiplano, Aguada Blanca National Reserve, Peru.

Cyril Ruoso / African elephant (Loxodonta africana) hanging upside down, during translocation by Kenya Wildlife Service, due to over population, Mwaluganje Reserve, Kenya. October 2006.

Sven Zacek / Four Roe deer (Capreolus capreolus) bucks with velvet antlers and the shadow of a fifth buck, Tartumaa, Estonia, March.

Roger Powell / Small flock of Brent Geese (Branta bernicla) in flight..Texel Island, The Netherlands, Europe

Staffan Widstrand / Kittiwakes (Rissa tridactyla) nesting on wall of house, Vardo town, Varanger Peninsula, Norway, March.

Andy Rouse / Magnificent frigate bird (Fregata magnificens) group flying against cloudy sky, Galapagos.

George Sanker / American buffalo (Bison bison) with group of calves, Yellowstone National Park, Wyoming, USA. May.

Staffan Widstrand / Wild Wonders of China / Crested myna starlings (Arcidotheres cristatellus) sitting on domestic waterbuffaloes (Bubalus domesticus), Poyang Ho Lake, Jiangxi province, China

Espen Bergersen / Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway. July

Chris & Monique Fallows / Long-beaked common dolphin (Delphinus capensis) school near the water?s surface, False Bay, South Africa

Enrique Lopez-Tapia / Hamer women in traditional clothing at the Jumping of the bulls ceremony. Ethiopia, November 2014

Denis-Huot / African elephant (Loxodonta africana) herd eating, Masai-Mara Game Reserve, Kenya

Ole Jorgen Liodden / Galapagos penguin (Spheniscus mendiculus) group underwater, Galapagos. Endemic.

Erlend Haarberg / Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.

Alex Mustard / Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea, close to Little Brother Island. The Brothers Islands, Egy...

Kristel Richard / Band of wild Mongolian horses running on plains at the foot of Dungurukh Uul mountain, near the border with China and Kazakhstan, Bayan-Olgiy aymag, Mongolia. September.

Jussi Murtosaari / Goldeneye (Bucephala clangula) with chicks swimming behind, Helsinki, Uusimaa, Finland, May.

Andy Rouse / Chinstrap penguin (Pygoscelis antarcticus) group on blue iceberg, Antarctica

Constantinos Petrinos / Bigfin reef squid (Sepioteuthis lessoniana) schooling parallel to one another. Lembeh Strait, North Sulawesi, Indonesia.