Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male displaying at display post, Malanda, Queensland, Australia

Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male displaying at display post, Malanda, Queensland, Australia

Martin Willis / Frilled Monarch (Arses telescophthalmus) male, Kutini-Payamu National Park, Queensland, Australia

Martin Willis / Scarlet Macaw (Ara macao) pair preening, Costa Rica

Martin Willis / Empress Brilliant (Heliodoxa imperatrix) and Buff-tailed Coronet (Boissonneaua flavescens) hummingbirds fighting, Andes, Ecuador

Martin Willis / Splendid Fairywren (Malurus splendens) female and male, Queensland, Australia

Martin Willis / Rainbow Lorikeet (Trichoglossus haematodus) pair preening during courtship, Magnetic Island, Queensland, Australia

Martin Willis / Budgerigar (Melopsittacus undulatus) father with begging chick at nest cavity, Queensland, Australia

Martin Willis / Rainbow Lorikeet (Trichoglossus haematodus), Malanda, Queensland, Australia

Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male displaying for female on vine lek, Malanda, Queensland, Australia

Martin Willis / Wompoo Fruit-Dove (Ptilinopus magnificus), Malanda, Queensland, Australia

Martin Willis / White-cheeked Honeyeater (Phylidonyris nigra), Cheyne Beach, Western Australia, Australia

Martin Willis / Grey-breasted Robin (Eopsaltria griseogularis) fluffing, Stirling Range National Park, Western Australia, Australia

Martin Willis / Victoria's Riflebird (Ptiloris victoriae) sub-adult male displaying at display post, Malanda, Queensland, Australia

Martin Willis / Splendid Fairywren (Malurus splendens) male, Busselton, Western Australia, Australia

Martin Willis / Sacred Kingfisher (Todirhamphus sanctus), Daintree River, Queensland, Australia

Martin Willis / Rufous Fantail (Rhipidura rufifrons), Malanda, Queensland, Australia

Martin Willis / Red-eared Firetail (Stagonopleura oculata), Busselton, Western Australia, Australia

Martin Willis / Papuan Pitta (Erythropitta macklotii), Kutini-Payamu National Park, Queensland, Australia

Martin Willis / Palm Cockatoo (Probosciger aterrimus), Kutini-Payamu National Park, Queensland, Australia

Martin Willis / Striated Grasswren (Amytornis striatus), Opalton, Queensland, Australia

Martin Willis / New Holland Honeyeater (Phylidonyris novaehollandiae), Cheyne Beach, Western Australia, Australia

Martin Willis / New Holland Honeyeater (Phylidonyris novaehollandiae) pair, Cheyne Beach, Western Australia, Australia

Martin Willis / Macleay's Honeyeater (Xanthotis macleayanus) pair, Malanda, Queensland, Australia

Martin Willis / Frilled Monarch (Arses telescophthalmus) male, Kutini-Payamu National Park, Queensland, Australia

Martin Willis / Carnaby's Black Cockatoo (Calyptorhynchus latirostris) feeding, Waychinicup National Park, Western Australia, Australia

Martin Willis / Baudin's Black Cockatoo (Zanda baudinii), Busselton, Western Australia, Australia

Martin Willis / Azure Kingfisher (Ceyx azureus), Daintree River, Queensland, Australia

Martin Willis / Black-mandibled Toucan (Ramphastos ambiguus) with predated egg at nest cavity, Costa Rica

Martin Willis / Splendid Leaf Frog (Agalychnis calcarifer), La Paz Waterfalls, Costa Rica

Martin Willis / Scarlet Macaw (Ara macao), Costa Rica

Martin Willis / Red-eyed Tree Frog (Agalychnis callidryas), La Paz Waterfalls, Costa Rica

Martin Willis / Plate-billed Mountain-Toucan (Andigena laminirostris), Andes, Ecuador

Martin Willis / Masked Flower-piercer (Diglossa cyanea), Andes, Ecuador

Martin Willis / Many-spotted Hummingbird (Taphrospilus hypostictus), Andes, Ecuador

Martin Willis / Many-banded Aracari (Pteroglossus pluricinctus), Amazon, Ecuador

Martin Willis / Many-banded Aracari (Pteroglossus pluricinctus) group, Amazon, Ecuador

Martin Willis / Long-tailed Sylph (Aglaiocercus kingi) hummingbird feeding on flower nectar, Andes, Ecuador

Martin Willis / Long-tailed Sylph (Aglaiocercus kingi) hummingbird, Andes, Ecuador

Martin Willis / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Martin Willis / Variable Mountain-gem (Lampornis castaneoventris) feeding on flower nectar, Costa Rica

Martin Willis / Golden Tanager (Tangara arthus), Andes, Ecuador

Martin Willis / Gliding Leaf Frog (Agalychnis spurrelli), La Paz Waterfalls, Costa Rica

Martin Willis / Chestnut-breasted Coronet (Boissonneaua matthewsii) feeding on flower nectar, Andes, Ecuador

Martin Willis / Buff-tailed Coronet (Boissonneaua flavescens) hummingbird feeding on flower nectar, Andes, Ecuador

Martin Willis / Andean Cock-of-the-rock (Rupicola peruvianus) male, Andes, Ecuador

Martin Willis / Yellow-rumped Munia (Lonchura flaviprymna) male, Kimberley, Western Australia, Australia

Martin Willis / Victoria's Riflebird (Ptiloris victoriae) male in courtship display, Atherton Tableland, Queensland, Australia. Sequence 3 of 3