Nick Upton / Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK,...
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Nick Upton / White stork (Ciconia ciconia) with three chicks in nest at top of an Oak tree, gaping in hot sunshine with one exercising its wings, Knepp Estate, Sussex, UK, June.
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Nick Upton / Heath sand wasp (Ammophila pubescens) excavating a nest burrow in a bare sandy patch of heathland, flying back and forth, carrying away balls of sand between its mandibles and its raised front legs, D...
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Nick Upton / Common carder bumblebee (Bombus pascuorum) nectaring on a Dandelion (Taraxacum officinale) flowerhead among many Common daisies (Bellis perennis) on a lawn left unmown to allow wild flowers to bloom t...
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Nick Upton / Roe deer (Capreolus capreolus) buck running past a garden shed at night, Wiltshire, UK, April. Property released.
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Nick Upton / Greylag goose (Anser anser) landing with its wings and head raised to chase off another from its territory in a marshland pool, Gloucestershire, UK, January.
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Nick Upton / Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December.
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Nick Upton / Common bee fly (Bombylius major) nectaring on a Greater periwinkle flower (Vinca major), Wiltshire garden, UK, April.
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Nick Upton / Common carder bumblebee (Bombus pascuorum) nectaring on a Yellow archangel (Lamiastrum galeobdolon) flower in a garden, Wiltshire, UK, April
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Nick Upton / Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire,...
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Nick Upton / White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April.
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Nick Upton / Seven-spot ladybird (Coccinnella septempunctata) just emerged from hibernation, sunning on a Honeysuckle leaf with buildings in the background, Wiltshire garden, UK, April.
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Nick Upton / Honey bee (Apis mellifera) nectaring on a Crab apple (Malus sylvestnis) flower in a garden, Wiltshire, UK, April.
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Nick Upton / Garden snail (Cornu aspersum / Helix aspersa) crawling over an oak sleeper retaining a garden lawn with buildings and a greenhouse in the background, Wiltshire, UK, April.
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Nick Upton / Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...
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Nick Upton / Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...
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Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...
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Nick Upton / Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...
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Nick Upton / White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.
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Nick Upton / Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...
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Nick Upton / Recently hatched White stork (Ciconia ciconia) chicks begging for food in their nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cots...
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Nick Upton / Pair of White storks (Ciconia ciconia) mating on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for several hundreds...
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Nick Upton / Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
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Nick Upton / Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
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Nick Upton / White stork (Ciconia ciconia) pair mating on their nest at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White storks nesting in the UK for several hundreds of y...
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Nick Upton / White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White s...
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Nick Upton / White stork (Ciconia ciconia) male landing with nest material and joining his mate on their nest in an Oak tree at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of...
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Nick Upton / View towards Naranjo de Bulnes (Picu Urriellu) a distinctive 2519m limestone peak, Picos de Europa mountains, viewed from Arenas de Cabrales at sunset, Asturias, Spain, August 2016.
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Nick Upton / Mediterranean sea holly / Pyrenean eryngo (Eryngium bourgatii) flowering on montane pastureland, Covadonga, Picos de Europa, Asturias, Spain, August.
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Nick Upton / Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August.
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Nick Upton / Cornish heath (Erica vagans) clump flowering on montane pastureland, above the Lakes of Covadonga, at 1300m, Picos de Europa, Asturias, Spain, August.
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Nick Upton / Red mason bees (Osmia bicornis) entering and leaving nest hole drilled into a log within an insect hotel, Gloucestershire, England, UK, April.
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Nick Upton / Ivy bee (Colletes hederae) mating ball, showing a mass of males clustered around a female recently emerged from her burrow, Wiltshire, England, UK, September.
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Nick Upton / Two Red fox (Vulpes vulpes) cubs playing near the entrance to their den at dusk, Somerset, England, UK, May.
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Nick Upton / European eel (Anguilla anguilla) elver migrating upstream, using an eel pass on the side of a weir, Somerset Levels, England, UK, July.
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Nick Upton / Male Eurasian beaver (Castor fiber) grazing at dusk, River Otter, Devon, England, UK, June.
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Nick Upton / Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.
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Nick Upton / Forest caterpillar hunter (Calosoma sycophanta) feeding on a Gypsy moth caterpillar (Lymantria dispar) it has killed in a Holm oak tree, Bacu Goloritze ravine, Baunei, Sardinia, Italy, May.
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Nick Upton / Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June.
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Nick Upton / Feral horse (Equus caballus) walking on a remote sandy beach, near Arta, Mallorca, Spain, August.
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Nick Upton / Female House spider (Tegenaria sp.) at the mouth of her tubular silk retreat in an old stone wall, Somerset, UK, September.