Simon Littlejohn / Two European otters (Lutra lutra) hunting in the River Stour, Dorset, England, UK, May.

Simon Littlejohn / Juvenile Lammergeier vulture (Gypaetus barbatus) in flight, mobbed by a raven, Grand Bargy cirque, Haute Savoie, France, October.

Simon Littlejohn / Bonelli's eagle chicks (Aquila fasciata) in nest pecking each other, aged one week, Andalusia , Spain, May.

Simon Littlejohn / Pair of Griffon vultures (Gyps fulvus) mating, Andalusia, Spain, June.

Simon Littlejohn / Honey bee (Apis) nectaring from a flowering Eucalyptus tree, Andalusia, Spain, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) feeding on a Guanaco (Lama guanacoe) carcass, Torres del Paine National Park, Patagonia, Chile, July.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) dragging the carcass of a young Guanaco (Lama guanicoe), Torres del Paine National Park, Patagonia, Chile, July.

Simon Littlejohn / Close-up of a female Patagonian puma (Puma concolor patagonica) feeding on a Guanaco (Lama guanicoe) carcass, filmed with a remote camera, Torres del Paine National Park, Patagonia, Chile, May.

Simon Littlejohn / Flock of Griffon vultures (Gyps fulvus) feeding, Andalusia, Spain, May.

Simon Littlejohn / Griffon vulture (Gyps fulvus) regurgitating food for its chick, chick struggles to eat, Andalusia, Spain, June.

Simon Littlejohn / Mute swans (Cygnus olor) hissing and lunging at a European otter (Lutra lutra), River Stour, Dorset, England, UK, June.

Simon Littlejohn / Family of European otters (Lutra lutra) feeding and playing amongst Whitewater crowfoot (Ranunculus aquatilis) flowers, River Stour, Dorset, England, UK, June.

Simon Littlejohn / Close-up of a female European otter (Lutra lutra) and two juveniles feeding in a river, River Stour, Dorset, England, UK, June.

Simon Littlejohn / Female European otter (Lutra lutra) and two large pups playing and feeding in mid river, River Stour, Dorset, England, UK, June.

Simon Littlejohn / Adult female European otter (Lutra lutra) standing in shallow water by river bank with her two pups (approximately 12 weeks old), one pup is restingo n her back until she dives under the water and the...

Simon Littlejohn / Adult female European otter (Lutra lutra) fishing with her two pups (approximately 12 weeks old) following her as she clambers out on to riverbank with a small fish to feed them, River Stour, Dorset,...

Simon Littlejohn / Two European otter (Lutra lutra) pups (approximately 12 weeks old) swimming beside their mother who is diving for food, River Stour, Dorset, UK, October.

Simon Littlejohn / Two European otter (Lutra lutra) pups (approximately 12 weeks old) sitting in the shallows waiting for their mother to return before they bothturn and dive into the water, River Stour, Dorset, UK, Oct...

Simon Littlejohn / Red deer (Cervus elaphus) stags rutting, with one stag urinating as they fight, Bushy Park, London, UK, September.

Simon Littlejohn / Slow worm (Anguis fragilis) resting in a patch of dry heather and bracken before slithering away under a heather bush, Wareham Forest, Dorset, UK, July.

Simon Littlejohn / Stoat (Mustela erminea) running and hunting, with winter coat, Haute-Savoie, France, October.

Simon Littlejohn / Female Patagonian puma (Puma concolor Patagonia) sheltering from a blizzard in bushes,Torres del Paine National Park, Patagonia, Chile, July.

Simon Littlejohn / Lammergeier vulture (Gypaetus barbatus) soaring, Monte Perdido y Ordesa National Park, Aragon, Spain, September.

Simon Littlejohn / Three juvenile Spanish imperial eagles (Aquila adalberti) in nest, one flapping its wings, Sierra de Andujar National Park, Andalusia, Spain, July.

Simon Littlejohn / Two Patagonian puma cubs (Puma concolor patagonica) playing, Torres del Paine National Park, Chile, April.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) walking through tall grass, with cub running in front of her, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Violet dropwing dragonfly (Trithemis annulata) taking off and landing on a stick, Sierra de Andujar Natural Park, Andalusia, Spain, August.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) walking through tall grass with two cubs, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Bonelli's eagle (Aquila fasciata) chick defecating out of nest, Andalusia, Spain, May.

Simon Littlejohn / Bonelli's eagle (Aquila fasciata) feeding in nest whilst brooding two chicks, takes off, Andalusia, Spain, April.

Simon Littlejohn / Juvenile Griffon vulture (Gyps fulvus) begging for food from parent, parent regurgitates, Grazalema National Park, Andalusia, Spain, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) stalking, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Wide-angle shot of a Patagonian puma (Puma concolor patagonia) sitting on a hillside, with mountains in the background, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) stretching and sharpening claws in snow covered landscape, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) resting on hillside with two cubs, Torres del Paine National Park, Chile, July.

Simon Littlejohn / Common caracara (Caracara plancus) scavenging on a Guanco (Lama guanacoe) carcass, with a South American grey fox (Lycalopex griseus) in the background, Torres del Paine National Park, Chile, South Am...

Simon Littlejohn / Darwin's rhea (Rhea pennata) preening its wing feathers, Torres del Paine National Park, Chile, July.

Simon Littlejohn / Male juvenile Patagonian puma (Puma concolor patagonia) grooming its claws, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Male juvenile Patagonian puma (Puma concolor patagonia) scratching, before walking off, Torres del Paine NP, Chile, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) and juvenile male feeding on a Guanaco (Lama guanacoe) carcass, Torres del Paine National Park, Patagonia, Chile, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) and juvenile male feeding on a Guanaco (Lama guanacoe) carcass, Torres del Paine National Park, Patagonia, Chile, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) and four cubs walking along the shore of a lake, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Adult Iberian lynx (Lynx pardel) resting on a boulder and looking around, Sierra de Andujar National Park, Andalusia, Spain, May.

Simon Littlejohn / Bonelli's eagle (Aquila fasciata) feeding chick in the nest, Andalusia, Spain, May.

Simon Littlejohn / Two Bonelli's eagle (Aquila fasciata) chicks in the nest, pecking each other, Andalusia, Spain, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) resting in bushes, grooming claws, Torres del Paine National Park, Patagonia, Chile, June.

Simon Littlejohn / Close-up of a female Patagonian puma (Puma concolor patagonica) feeding on a Guanaco (Lama guanicoe) carcass, filmed with a remote camera, Torres del Paine National Park, Patagonia, Chile, May.

Simon Littlejohn / Close-up of a female Patagonian puma (Puma concolor patagonica) feeding on a Guanaco (Lama guanicoe) carcass, filmed with a remote camera, Torres del Paine National Park, Patagonia, Chile, May.