Neil Challis / Infant Long-tailed macaque (Macaca fascicularis) suckling and cuddling with mother, before being snatched by another adult, Central Thailand, September.
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Vishal Jadhav / Coppersmith barbet (Psilopogon haemacephalus) hanging upside down on a fruiting tree, filling its beak with berries to take back to the nest, then hops away, Maharashtra India, April.
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Marie Read / Eastern bluebird (Sialia sialis) male feeding on Winterberries (Ilex verticillata) before flying off, Freeville, New York, USA, January.
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John Waters / Golden plover (Pluvialis apricaria) flock taking off and flying over saltmarsh, Steart Marshes WWT Reserve, Somerset, UK, November.
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Vishal Jadhav / Red-breasted flycatcher (Ficedula parva) male hunting flying insects from a rock, Maharashtra, India, February.
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Vishal Jadhav / Spotted owlet (Athene brama) female looking out of nest in rocks, males appears then flies off, Maharashtra, India, January.
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Oliver Wright / Marbled white butterfly (Melanargia galathea) sequence showing landing and taking off from a Greater knapweed (Centaurea scabiosa) flower. Yorkshire, UK. Digital composite.
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Markus Varesvuo / Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October.
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Mateusz Piesiak / Two Common snipe (Gallinago gallinago) foraging in mud in thick fog, Lower Silesia, Poland. Emotion'Ailes Photography Competition 2021, Category Winner - Bird in all its states.
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Terry Whittaker / White-throated dipper (Cinclus cinclus) returning to newly-fledged chick to feed it before flying away, River Mersey, Greater Manchester, UK.
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Kim Taylor / Male Cockchafer beetle (Melolontha melolontha) unfolding its wings before taking off to fly away, Surrey, UK, May.
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Richard Hopkins / Slow motion shot of Short-eared owl (Asio flameus) taking off from the ground and flying out of frame, Gloucestershire, UK, March.
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Richard Hopkins / Slow motion shot of Short-eared owl (Asio flameus) flying over grassland searching for prey, before it dives into the grass and then flies away, Gloucestershire, UK, March.
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John Chan / Calliope hummingbird (Stellula calliope) chasing insects before returning to its perch, Southern California, USA, April.
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John Chan / Monarch butterfly (Danaus plexippus) taking flight from a Pride of madeira (Echium candicans) after contact with a Honey bee (Apis mellifera), Southern California, USA, April.
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John Chan / Elegant terns (Thalasseus elegans) gathering at mussel bed during breeding season, one hovering tern has a mussel clamped to its foot, a common mishap caused by stepping on an open mussel, Bolsa Chica...
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Guy Edwardes / Long-billed curlew (Numenius americanus) taking off along shoreline. Fort De Soto Park, Florida, USA. April.
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Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male from branch, flies off, Bedfordshire, England, UK, May.
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Brian Bevan / Male European cuckoo (Cuculus canorus) trying to mate with a decoy female, used as part of a scientific study, Bedfordshire, England, UK, May.
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Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male, before flying off in response to a female, Bedfordshire, England, UK, May.
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Morley Read / Slow motion shot of a Rufous tailed hummingbird (Amazilia tzacatl) drinking nectar from a Heliconia flower, Mindo, Ecuador.
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Jack Dykinga / Yellow-headed blackbird (Xanthocephalus xanthocephalus) flock murmuration as they take off from the marsh at dawn, Whitewater Draw, Arizona State Game and Fish Reserve, USA. January.
Markus Varesvuo / White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August.
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Andres M. Dominguez / Common firecrest (Regulus ignicapilla) taking off, Sierra de Grazalema Natural Park. Southern Spain. September
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Andres M. Dominguez / Common redstart (Phoenicurus phoenicurus) male taking off, Sierra de Grazalema Natural Park. Southern Spain. September
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Morley Read / Slow motion clip of a large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador.
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Morley Read / Large group of Butterflies (Pieridae) taking flight after puddling, absorbing nutrients or minerals from damp soil, Orellana Province, Amazon rainforest, Ecuador.
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Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Camera attached to helmet, Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.
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Terry Whittaker / Man being lowered down a cliff to collect seabird eggs, including those of Common guillemots (Uria aalge). Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland, May.
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John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019
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Jack Dykinga / Flocks of Mallard ducks (Anas platyrhynchos) taking off in tree-lined wetlands. Bosque del Apache National Wildlife Refuge, New Mexico, USA.
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Tony Heald / Glaucous Gull (Larus hyperboreus) taking off from ice, Svalbard Norway, July.
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John Chan / Female Anna's hummingbird (Calypte anna) incubating eggs, departs from nest momentarily to feed, Southern California, USA, March.
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John Chan / Female Anna's hummingbird (Calypte anna) incubating eggs, departs from nest momentarily to feed, Southern California, USA, March.
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Michael Hutchinson / Marsh fritillary butterfly (Euphydryas aurinia) nectaring from a Buttercup (Ranunculus) before taking off, Devon, England, UK, June.
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John Chan / Elegant tern (Thalasseus elegans) with a broken wing, attempting to take off, Southern California, USA, April.
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Kerstin Hinze / Painted lady (Vanessa cardui) nectaring, Bavarian Forest National Park, Germany, June.
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Philippe Clement / Common raven (Corvus corax) taking off from the top of a sea cliff, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
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Dave Bevan / Slow motion clip of a pair of Common swifts (Apus apus) approaching and entering a nest hole in limestone cliff, Norfolk, England, UK, May.
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Juergen Freund / Colony of Little red flying foxes (Pteropus scapulatus) leaving roost site, Atherton Tablelands, Queensland, Australia.
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John Chan / Honey bee (Apis mellifera) gathering pollen from a Sea fig (Carpobrotus chilensis), Southern California, USA, April.
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Konrad Wothe / Chub (Squalius cephalus) trying to catch a Banded demoiselle (Calopteryx splendens), Bavaria, Germany, June.
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John Waters / Slow motion clip of a Brown trout (Salmo trutta) predating an adult Mayfly (Ephemoptera) struggling to take off from water, River Kennet, Hungerford, Berkshire, England, UK, June.
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Staffan Widstrand / Wild Wonders of China / Crested ibis (Nipponia nippon) flying, Yangxian Nature Reserve, Shaanxi, China, September.
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Dave Bevan / Slow motion clip of a Tawny owl (Strix aluco) leaving nest site in hollow tree, Carmarthenshire, Wales, UK, April.
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John Waters / Wide-angle shot of wetland landscape at sunset, with Lapwings (Vanellus vanellus) flying to roost, Ham Wall RSPB Reserve, Somerset Levels, England, UK, February.
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Michael Hutchinson / Slow motion clip of a Common carder bumblebee (Bombus pascuorum) feeding from a Viper's bugloss (Echium vulgare) flower, Bristol, England, UK, September.