Markus Varesvuo / Great Grey Owl (Strix nebulosa) hunting over snow, Kuhmo Finland, March.
Markus Varesvuo / Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March.
Markus Varesvuo / Black Woodpecker (Dryocopus martius) male in flight, Finland, February.
Markus Varesvuo / Sparrowhawk (Accipiter nisus) adult female, Norway, October.
Espen Bergersen / Northern fulmar (Fulmarus glacialis) in front of the Seljalandsfoss waterfall, Iceland. May
Espen Bergersen / Common Loon (Gavia immer) flapping wings in water, Iceland, May.
Andres M. Dominguez / Black-tailed godwit (Limosa limosa) Danube Delta, Romania. July
Andres M. Dominguez / Common firecrest (Regulus ignicapilla) taking off, Sierra de Grazalema Natural Park. Southern Spain. September
Andres M. Dominguez / Common redstart (Phoenicurus phoenicurus) male taking off, Sierra de Grazalema Natural Park. Southern Spain. September
Hermann Brehm / Cuckoo (Cuculus canorus) in flight, Germany, April. May
Oscar Dewhurst / Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April.
Guy Edwardes / Barn owl (Tyto alba) in flight about to land on post in morning light, Hampshire, England, UK. July. Controlled conditions.
George Sanker / Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July. This island is disputed between USA and Canada.
Piotr Naskrecki / Termite (Macrotermes mossambicus) winged individual on pile of wings shed by other termites, with winged male Carpenter ants (Camponotus). Gorongosa National Park, Mozambique. Highly commended in the...
John Chan / Great horned owl (Bubo virginianus) chicks stretching and flapping their wings, Southern California, USA, May.
John Cancalosi / Great horned owl (Bubo virginianus) chick stretching its wings in nest in a Saguaro (Carnegiea gigantea), is groomed by its parent, Sonoran Desert, Arizona, USA, May.
Andy Rouse / Mute swan (Cygnus olor) stretching wings on water. In morning light, Wales, UK. February.
Oscar Dewhurst / Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September
Phil Savoie / Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May.
Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.
Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Marie Read / Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April.
Marie Read / Ruby-throated Hummingbird (Archilocus colubris), male flying in to feed from cardinal flowers (Lobelia cardinalis) New York, USA. August.
Marie Read / Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
Bernard Castelein / Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Bernard Castelein / Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India
Bernard Castelein / Intermediate egret (Egrettta intermedia), Keoladeo NP, Bharatpur, India
Andy Rouse / Barn owl (Tyto alba) hunting with sunset light on wings, UK. March.
Alex Hyde / Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November.
David Pattyn / Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July
Staffan Widstrand / Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January.