Marie Read / American bittern (Botaurus lentiginosus) calling / booming in a cattail marsh to mark territory, Salt Alkaline Lake, Kidder County, North Dakota, USA, June.
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Vishal Jadhav / Exhausted Short-eared owl (Asio flammeus) surrounded by House crow (Corvus splendens) group at dawn, one crow dives down to attack, Maharashtra, India, April.
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Vishal Jadhav / Indian jackal (Canis aureus indicus) pair howling, Maharashtra, India, September.
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Brian Bevan / Yellowhammer (Emberiza citronella) male singing from bramble then flies away, Bedfordshire, UK, June.
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Terry Whittaker / White-throated dipper (Cinclus cinclus) singing on old bicycle wheel, River Mersey, Greater Manchester, UK.
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Doug Gimesy / Sub-Antarctic / Brown skua (Stercorarius antarcticus) flying away with egg stolen from a Royal penguin colony (Eudyptes schlegeli), Macquarie Island, Australian Territory. December.
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Claudio Contreras / Laysan albatross (Phoebastria immutabilis) calling mate,Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
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Ismaele Tortella / Eurasian otter (Lutra lutra) swimming with snow on its back, calling, Norway
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Will Burrard-Lucas / A mating pair of leopards (Panthera pardus) caught in front of a camera trap at night, Laikipia Wilderness Camp, Kenya. Communication Arts Annual 62 Competition 2021 Winner Book Category; Sony World P...
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Edwin Giesbers / Grey Wolf (Canis lupus), snarling protectively over carcass, Biotoppark Anholt, Germany, May. Captive.
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Alex Mustard / Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home, a Haddon's sea anemone (Stichoda...
01654192
Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male from branch, flies off, Bedfordshire, England, UK, May.
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Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male, before flying off in response to a female, Bedfordshire, England, UK, May.
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Konrad Wothe / Macgregor's bowerbird (Amblyornis macgregoriae) singing in rainforest, Papua New Guinea.
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Marie Read / Red-winged Blackbird (Agelaius phoenicus) male singing from snow-covered red maple (Acer rubrum) in flower, Ithaca, New York, USA. April 2020.
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Danny Green / Red deer stag (Cervus elaphus) roaring. Bradgate Park, Leicestershire, UK, October.
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Erlend Haarberg / Black grouse (Lyrurus tetrix), male calling early in the morning, Norway. May.
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Markus Varesvuo / Red-throated Diver (Gavia stellata) calling, Iceland, June.
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Felis Images / Asiatic lion (Panthera leo persica), female and cubs in morning light, cub biting mother's leg. Gir National Park, Gujarat, India. Photo?hillip Ross/Felis Images
David Gallan / Cameratrap video of a male Superb lyrebird (Menura novaehollandiae) courting female on display mound, Australia, July.
01620559
Klein & Hubert / Black / Melanistic Leopard (Panthera pardus) growling. Controlled conditions
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Klein & Hubert / Grey wolf (Canis lupus) female and two cubs, one howling. Controlled conditions
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Klein & Hubert / Wolf (Canis lupus) puppy, age one month, Rocky Mountains, USA. Controlled conditions.
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Stefan Christmann / Adelie penguin (Pygoscelis adeliae) calling during courtship, Atka Bay, Queen Maud Land, Antarctica, December.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August. Sequence 1/2.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), group of three males incubating, one with begging chick. Atka Bay, Antarctica. August.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
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Guy Edwardes / Marsh Frog (Pelophylax ridibundus) vocalising with bokeh effect in the background, Danube Wetlands, Bulgaria
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Guy Edwardes / Marsh Frog (Pelophylax ridibundus) vocalising with bokeh effect in the background, Danube Wetlands, Bulgaria
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David Perpinan / Male Mediterranean tree frog (Hyla meridionalis) calling, showing vocal sac inflating, Barcelona, Spain, May. Filmed at night with an infrared camera.
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Paul Hobson / Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May.
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Ann & Steve Toon / Lion (Panthera leo) male snarling at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September
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Philippe Clement / Territorial Common raven (Corvus corax) on cliff top, tearing out vegetation, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
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Philippe Clement / Common raven (Corvus corax) calling from top of sea cliff, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
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Dave Bevan / Male Pied flycatcher (Ficedula hypoleuca) calling to establish territory and attract a mate, Carmarthenshire, Wales, UK, May.
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Paul Williams / Lion (Panthera leo) close up of teeth while its snarling, captive.
01610843
Hermann Brehm / Nightingale (Luscinia megarhynchos ) singing, Germany, April.
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Konrad Wothe / Male Blackbird (Turdus merula) singing, Bavaria, Germany, April.
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Guy Edwardes / Little bee-eater (Merops pusillus) vocalising whilst perched on branch, Khwai, Botswana.
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Konrad Wothe / Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June.
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Dave Bevan / Robin (Erithacus rubecula) singing, Carmarthenshire, Wales, UK, February.
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Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.
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Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.
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Phil Savoie / Swift (Apus apus) flock screaming in flight against blue sky and clouds, Monmouthshire Wales, UK, July.
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Phil Savoie / Swift (Apus apus) flock screaming in flight against blue sky and clouds, Monmouthshire, Wales, UK, July.
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David Perpinan / Oystercatcher (Haematopus ostralegus) incubating eggs on nest, alarm calling as a predator flies overhead, Scotland,UK, June.
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Tui De Roy / Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
