Owen Newman / Male Leopard resting in tree {Panthera pardus} S Luangwa NP Zambia

Klein & Hubert / Cape buffalo (Syncerus caffer) with Yellow-billed oxpckers (Buphagus africanus) drinking at waterhole in dry season, South Luangwa National Park, Zambia, Africa.

Ben Cranke / African fish eagle (Haliaeetus vocifer) pressurises a Saddle-billed stork (Ephippiorhynchus senegalensis) into dropping a freshly caught fish, so that it can be stolen for a meal. Liuwa Plain National...

Ben Cranke / African fish eagle (Haliaeetus vocifer) attacking a female Saddle-billed Stork (Ephippiorhyncus senegalensis )trying to steal a fish it has just caught. Liuwa Plains National Park, Zambia

Klein & Hubert / African elephant (Loxodonta africana) herd crossing the Luangwa river, Luangwa National Park, Zambia

Nick Garbutt / Adult male lion (Panthera leo) standing on the banks of the Luangwa River, South Lunangwa NP. Zambia.

Klein & Hubert / Hippo (Hippopotamus amphibius) walking to the Luangwa River at sunrise, Luangwa National Park, Zambia.

Klein & Hubert / Young Hippo (Hippopotamus amphibius) trotting to the Luangwa River at sunrise, Luangwa National Park, Zambia.

Neil Aldridge / White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia.

Eric Baccega / Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia

Eric Baccega / Coalition of two African male lions (Panthera leo) resting on the banks of the Luangwa river, dry season, South Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) opening mouth in river - detail upper jaw, nostrils and lips, Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) female and young entering water in the Luangwa river- portrait, Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) in river - portrait, Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) with Red-bellied oxpeckers (Buphagus erythrorhynchus) on his head in river, Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) charging in Luangwa river, close up of mouth and teeth, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) standing in water the Luangwa River at sunrise, Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) walking to the Luangwa River at sunrise, Luangwa National Park, Zambia

Klein & Hubert / Hippopotamus (Hippopotamus amphibius) silhouetted, walking to the Luangwa River at dawn, Luangwa National Park, Zambia

Neil Aldridge / Lion cub (Panthera leo) calling to its mother from the raised deck of a tourist lodge in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable species.

Neil Aldridge / Lion cub (Panthera leo) playfully attacking its mother as she drags her puku antelope prey across burnt Busanga Plains, Kafue National Park, Zambia. Vulnerable

Neil Aldridge / African lion (Panthera leo) male lying in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable

Nick Garbutt / Leopard (Panthera pardus) female drinking at night. South Luangwa National Park, Zambia.

Will Burrard-Lucas / Spotted hyena (Crocuta crocuta) at night, taken with remote camera. Liuwa Plain National Park, Zambia. December

Will Burrard-Lucas / Spotted hyena (Crocuta crocuta) low angle shot of group taken with remote camera. Liuwa Plain National Park, Zambia. May

Luke Massey / Cape pangolin (Smutsia temminckii) foraging at night. South Luangwa National Park, Zambia.

Andy Rouse / African leopard (Panthera pardus) South Luangwa, Zambia.

Felicity Lanchester,Felicity Lanchester / Fork-tailed drongo (Dicrurus adsimilis) catching insects fleeing from a grassland fire, South Luangwa National Park, Zambia.

Nick Garbutt / Adult bull African Elephant (Loxononta africana) trunk reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October

Nick Garbutt / Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October

Nick Garbutt / Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November

Nick Garbutt / Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched togethe...

Nick Garbutt / Juvenile African elephant (Loxodonta africana) wanders through a tourist safari lodge, Mfuwe Lodge, South Luangwa NP, Zambia, October

Nick Garbutt / Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October

Andy Rouse / African Lion (Panthera leo) adult looking from behind tree, half hidden. South Luangwa, Zambia, Africa.

Andy Rouse / African lion (Panthera leo) playing with Burchells zebra (Equus quagga burchelli) originally chased but not serious, Luangwa Valley, Zambia, Africa (non-ex)

Andy Rouse / African lion (Panthera leo) hunting Buffalo, South Luangwa, Zambia (non-ex)

Andy Rouse / Chimpanzee (Pan troglodytes) mother with youngster, captive, Chimfunshi Orphanage, Zambia. (non-ex).

Andy Rouse / African elephant (Loxodonta africana) breeding herd crossing river at sunset, South Luangwa, Zambia (non-ex)

Andy Rouse / African elephant (Loxidonta africana) female and calf in breeding herd, South Luangwa NP, Zambia

Andy Rouse / African elephant (Loxidonta africana) female and calf in breeding herd, South Luangwa NP, Zambia

Mark Carwardine / A cloud of Straw-coloured fruit bats (Eidolon helvum) with an immature African fish eagle (Haliaeetus vocifer) flying amongst them, Kasanka National Park, Zambia, Africa

Mark Carwardine / A cloud of Straw-coloured fruit bats (Eidolon helvum) flying around daytime roost with others still hanging from branches, Kasanka National Park, Zambia, Africa

Mark Carwardine / Straw-coloured fruit bat (Eidolon helvum) flying at night, Kasanka National Park, Zambia, Africa

Mark Carwardine / Straw-coloured fruit bats (Eidolon helvum) flying over daytime roost Kasanka National Park, Zambia, Africa

TJ Rich / African lion {Panthera leo} pride on alert, South Luangwa NP, Zambia

TJ Rich / Hippopotamus in Luangwa river at sunrise, South Luangwa National Park, Zambia

Martha Holmes / Cape buffalo {Syncerus caffer caffer} large herd drinking in Luangwa river, Zambia.